The Snow Hill Island Formation is an Early Maastrichtian geologic formation found on James Ross Island, James Ross Island group, Antarctica.[1] Remains of a paravian theropod Imperobator antarcticus[2] have been recovered from it, as well as the elasmarian ornithopods Trinisaura santamartaensis, "Biscoveosaurus" and Morrosaurus antarcticus, the ankylosaurian Antarctopelta oliveroi, and the shark Notidanodon sp. Alongside these described genera are also the remains of indeterminate elasmosaurids,[3] lithostrotian titanosaurs and an indeterminate pterosaur.[4]
In the Herbert Sound Member of the Snow Hill Island Formation, bivalves, ammonites, and fish were found.[5]
Paleofauna
Invertebrates
Corals
Arthropoda
| Genus
|
Species
|
Presence
|
Material
|
Notes
|
Images
|
| Angarestia
|
A. australensis
|
East side of Leal Bluff
|
Specimens[7]
|
A Glypheidae lobster
|
|
| Cristafrons
|
C. praescientis
|
East side of Leal Bluff
|
Specimens[7]
|
A Raninidae crab
|
|
| Hadrocarcinus
|
H. wrighti
|
East side of Leal Bluff
|
Specimens[7]
|
A Necrocarcinidae crab
|
|
| Hoploparia
|
H. stokesi
|
East side of Leal Bluff; Humps Island
|
Specimens[7]
|
A Clawed lobster
|
|
Mollusks
Vertebrates
Fish
| Genus
|
Species
|
Presence
|
Material
|
Notes
|
Images
|
| Albuliformes
|
Indeterminate
|
Santa Marta Cove
|
Isolated vertebra, IAA-IRJ2000-24[10]
|
|
|
| Antarctichthys
|
A. longipectoralis
|
Santa Marta Cove
|
MN 7838-V, single and incomplete specimen[11]
|
A Dercetidae fish
|
|
| Apateodus
|
cf. A. sp.
|
Santa Marta Cove
|
teeth[10]
|
Alepisauriformes fish
|
|
| Callorhinchus
|
C. sp.
|
Santa Marta Cove
|
teeth[4][12]
|
A Chimaera
|
|
| Cf. C. sp.
|
Cape Lamb
|
teeth[4][12]
|
| Centrophoroides
|
C. sp.
|
Santa Marta Cove
|
Teeth[4][12]
|
A Squalidae shark
|
|
| Chimaera
|
C. zangerli
|
Santa Marta Cove, James Ross Island.
|
Teeth[4]
|
A chimaera.
|
|
| Chlamydoselachus
|
C. thompsoni
|
Santa Marta Cove, James Ross Island.
|
Complete dentition.[10]
|
A frilled shark.
|
|
| Cretalamna
|
C. appendiculatta
|
Santa Marta Cove
|
Teeth.[12]
|
An Otodontidae shark
|
|
| Edaphodon
|
E. snowhillensis
|
Herbert Sound Member.[13]
|
MLP 13- I-26-1, Complete dentition.[13]
|
A large species of chimaera.
|
|
| Enchodus
|
E. sp.
|
Santa Marta Cove
|
IAA-IRJ2000-32, IAA-IRJ2000-33, teeth[10]
|
Enchodontidae fish
|
|
| Ichthyodectiformes
|
Indeterminate
|
Vega island
|
MLP 15-XI-7-11, body scale patch[4]
|
|
|
| Lamniformes
|
Indeterminate
|
Santa Marta Cove
|
(IAA-IRJ2000-9 to IAA-IRJ2000-10, BAS DJ.172.11, teeth[10]
|
|
|
| Notidanodon
|
N. dentatus
|
Santa Marta Cove; Vega Island
|
(MLP 95-IV-1), lateral tooth embedded in the hosting rock[4]
|
A cow shark.
|
|
| N. sp.
|
Santa Marta Cove[14]
|
|
| Paraorthacodus
|
P. antarcticus
|
Santa Marta Cove
|
IAA-IRJ2000-19; BAS DJ.136.2, teeth[10]
|
A Synechodontiform
|
|
| Rhinochimaeridae
|
Indeterminate
|
Santa Marta Cove
|
teeth[12]
|
|
|
| Scapanorhynchus
|
S. spp.
|
Santa Marta Cove
|
IAA-IRJ2000-11[10]
|
A Mitsukurinidae shark
|
|
| Cf. S. sp.
|
Santa Marta Cove
|
teeth[4]
|
| Sphenocephalidae
|
Indeterminate
|
False Island Point
|
DJ. 360.8, small, incomplete, and poorly-preserved specimen that lack its skull and the anterior part of the body.[4]
|
|
|
| Sphenodus
|
S. sp.
|
Santa Marta Cove
|
IAA-IRJ2000-17 and IAA-IRJ2000-18, teeth[10]
|
An Orthacodontid
|
|
| Squatina
|
S. cf.hassei
|
Santa Marta Cove
|
IAA-IRJ2000-10-IAA-IRJ2000-16[10]
|
An Angelshark
|
|
Pterosaurs
Dinosaurs
| Genus
|
Species
|
Presence
|
Material
|
Notes
|
Images
|
| Antarcticavis
|
A. capelambensis[16]
|
Cape Lambe Member, Vega Island.
|
A partial skeleton (SDSM 78147) consists of two thoracic vertebrae, the sternum keel, the right coracoid and shoulder blade, the sternal part of the left coracoid, the right upper arm, parts of the left upper arm, the proximal right ulna, the proximal left ulna and radius (articulated), the proximal right carpometacarpus, the proximal left carpometacarpus, the distal left carpometacarpus, the synsacrum, the right and left thighs, the proximal right tibiotarsus, the right and left distal tibiotarsus, and the proximal right tarsometatarsus.
|
An avialan of uncertain phylogenetic placement.
|
|
| Antarctopelta
|
A. oliveroi
|
Santa Marta Cove[17]
|
A partial skeleton (MLP 86-X-28-1) consists of three isolated teeth, part of the lower jaw with another tooth in situ, some other skull fragments, vertebrae of the neck, back, hips and tail, some shoulder and hip bones (scapula, ilium) a thigh bone (femur), foot and hand bones (five metapodials and two phalanges), and numerous pieces of armor.[17][14]
|
A parankylosaur
|
|
| "Biscoveosaurus"
|
Indeterminate
|
Cape Lamb Member.[18][19]
|
NHMUK PV R 36760, dentaries, teeth, a braincase, parts of the maxillae, forelimb elements, assorted vertebrae, and the pectoral girdle.
|
An ornithopod.
|
|
| Elasmaria
|
Indeterminate
|
Santa Marta Cove
|
Two isolated ungual phalanges (MLP 07-III-2-1 and MLP 07-III-2-2)[4]
|
|
|
| Indeterminate
|
Fortress Hill
|
limb, foot[20]
|
| Indeterminate
|
Vega Island
|
BMNH BAS R.2450[9]
|
| Gaviiformes
|
Indeterminate
|
Vega Island
|
MLP 98-I-10-47: incomplete tarsometatarsus; MLP 98-I-10-50: incomplete right tarsometatarsus; MLP 98-I-10-51: proximal end of left femur[21]
|
|
|
| Imperobator
|
I. antarcticus
|
Cape Lamb Member, Naze Peninsula.[22]
|
A single specimen that contains skull fragments possibly from the premaxilla, maxilla, and dentary as well as a caudal vertebra, teeth, and pedal elements, UCMP 276000.[20][23]
|
A potential Unenlagiinae paravian
|
|
| Lithostrotia
|
Indeterminate
|
Loma Verde, near Santa Marta Cove
|
11-II-20-1, incomplete middle caudal vertebra[24]
|
|
|
| Morrosaurus
|
M. antarcticus
|
Cape Lamb Member.[25]
|
Fragmentary right hind limb.[25]
|
An elasmarian ornithopod.
|
|
| Neornithes
|
Indeterminate
|
Vega Island
|
MLP 98-I-10-25, left tarsometatarsus[4]
|
|
|
| Ornithurae
|
Indeterminate
|
Cape Lamb
|
AMNH FARB 30920, thoracic vertebra[26]
|
|
|
| Theropoda
|
Indeterminate
|
Cape Lamb
|
MLP 15-I-7-2, isolated pedal phalax (digit III?)[4]
|
|
|
| Trinisaura
|
T. santamartaensis
|
Santa Marta Cove, in the lower levels of the formation.[27]
|
MLP08-III-1-1, disarticulated and partial skeleton that includes one incomplete dorsal vertebra, three sacral centra, seven caudal vertebrae; two incomplete dorsal rib shafts, one proximal haemal arch, incomplete right scapulocoracoid, incomplete right humerus, two metacarpals, both ilia, right pubis, right ischium, right femur, right distal tibia, incomplete metatarsal III, first phalanx of pedal digit III, two phalanges of pedal digit IV, and indeterminate fragments.
|
An elasmarian ornithopod.
|
|
Plesiosaurs
| Genus |
Species |
Presence
|
Material |
Notes |
Images
|
| Aristonectinae
|
Indeterminate
|
Santa Marta Cove
|
SGO.PV.6579, fragmentary postcranial skeleton preserving eight fragmentary caudal centra, two articular propodial heads (likely femora), an epipodial (likely a fibula), ventral portion of the right ilium, partial left pubis, and several rib portions[28]
|
|
|
| Elasmosauridae
|
Indeterminate
|
Monolithic Lake; Santa Marta Cove
|
"MLP 11-II-20-4, one cervical vertebra preserving a partial rib; MLP 86-X-28-3, two cervical vertebrae; MLP 86-X-28-(2–6), 10 posterior cervical vertebrae articulated with three pectoral vertebrae, part of two dorsal vertebrae, ribs, and indeterminate fragments."[29]
|
|
| Indeterminate
|
Santa Marta Cove
|
SGO.PV.6508, one isolated cervical vertebral centrum[28]
|
| Indeterminate
|
W Santa Marta Cove
|
MN 7820-V, four vertebral centra and fragments of another six centra[30]
|
| Indeterminate
|
Cape Lamb
|
MLP 98-I-10-20, an incomplete postcranial skeleton (juvenile)[31]
|
| Indeterminate
|
Vega Island
|
SDSM 78156 (a nearly complete, articulated torso, partial paddles, and neck and tail sections)[32]
|
| Plesiosauria
|
Indeterminate
|
The Naze
|
DJ.355.140, half of large water-worn vertebra, either anterior trunk or distal cervical vertebra[33]
|
|
| Plesiosauroidea
|
Indeterminate
|
The Naze; Cape Lamb
|
MLP 78-XI-1-2; MLP 78-XI-1-1, vertebrae and fragments[34]
|
|
| Vegasaurus
|
V. molyi
|
Cape Lamb member, Vega Island.
|
MLP 93-I-5-1, postcranial skeleton preserving a complete cervical region with 54 cervical vertebrae, three pectoral vertebrae, 17 dorsal vertebrae, three sacral vertebrae, anterior and medial caudal vertebrae, pectoral and pelvic girdles, forelimbs and hind limbs, ribs, and gastroliths.[35]
|
A plesiosaur.
|
| Weddellonectia
|
Indeterminate
|
Cape Lamb
|
MLP 15-I-7-6, partial skull comprising caudal half of pterygoids, basisphenoid, basioccipital, squamosal, exoccipitalopisthotic; cervical, dorsal, sacral and caudal centra; cervical and dorsal ribs, partially preserved coracoid, two partially preserved propodials.[36]
|
|
| Indeterminate
|
Cape Lamb
|
AMNH FARB 30877 is a largely articulated partial skeleton[37]
|
|
Mosasaurs
| Genus
|
Species
|
Presence
|
Material
|
Notes
|
Images
|
| Liodon
|
L. sp.
|
Cape Lamb
|
Teeth[4]
|
A Mosasaur
|
|
| Cf. L. sp.
|
Cape Lamb
|
Teeth[4]
|
| Mosasaurus
|
M. cf. M. lemonnieri
|
Cape Lamb
|
Teeth[4]
|
A Mosasaur
|
| Plioplatecarpus
|
cf. P. sp.
|
Cape Lamb
|
MLP 93-I-3-5, small tooth[31]
|
|
| Taniwhasaurus[38]
|
T. antarcticus
|
Santa Marta Cove; Al's Bird Site
|
IAA 2000-JR-FSM-1, containing a skull measuring 72 cm (28 in) long, teeth, some vertebrae, and rib fragments.
|
A tylosaur.
|
| cf. T. antarcticus
|
Al's Bird Site
|
MLP 93-I-3-7, a tooth crown[39]
|
| Tylosaurinae
|
Indeterminate
|
Vega Island
|
Juvenile skull[40]
|
|
Plants
|
|
|
|
Color key
|
|
Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
|
During the Campanian–Maastrichtian, the Antarctic Peninsula supported temperate, humid forests dominated by podocarps, araucarian conifers, and a diversifying group of angiosperms. Key angiosperm families included Nothofagaceae, Monimiaceae, Cunoniaceae, Proteaceae, Myrtaceae, Lauraceae, Atherospermataceae, Winteraceae, and extinct Sassafras-like forms.[41][42] Another important group is Asteraceae, with Dasyphyllum-like pollen, the oldest fossils ever found for the family.[43] At The Naze area, Podocarpaceae-Nothofagus rainforests thriving in lowland areas under cool-temperate, frost-free, and high-rainfall conditions, with understories rich in other angiosperms and ferns.[44] Smaller components, likely endemic, included lycophytes, bryophytes, Proteaceae, Liliaceae, Palmae or Microthyriaceae, and various herbaceous or shrubby dicotyledons.[44] Forests were structurally similar to modern Valdivian temperate forests. These plants were primarily found in riparian floodplains, with bryophyte-lycophyte rich swamps and Chlorophyta-rich lake margins, while influence of marine waters can be seen by Dinoflagellates. The climate was highly humid and seasonal, as evidenced by distinct growth rings in Agathoxylon fossil wood from Lachman Crags and The Naze, indicating a temperate but variable environment.[44] The flora reflects a transition from gymnosperm-dominated to mixed angiosperm-conifer ecosystems under polar greenhouse conditions.[41][44]
Pteridophytes, Lycophytes, and Bryophytes
| Taxa
|
Species
|
Locality
|
Material
|
Notes
|
Images
|
| Baculatisporites[42][44]
|
B. comaumensis
|
Santa Marta Cove; The Naze
|
Spore
|
Osmundaceae
|
|
| Biretisporites[44]
|
B. potoniaei
|
The Naze
|
Spore
|
Fern affinity
|
|
| Calamospora[44]
|
sp.
|
The Naze
|
Spore
|
Equisetales
|
|
| Camarozonosporites[44]
|
C. ohioensis
|
The Naze
|
Spore
|
Fern affinity
|
|
| Ceratosporites[44]
|
C. equalis
|
The Naze
|
Spore
|
Fern affinity
|
|
| Cyatheacidites[42][44]
|
C. annulatus
|
Santa Marta Cove; The Naze
|
Spore
|
Dicksoniaceae (Lophosoria)
|
|
| Cyatheacidites[44]
|
C. archangelskii
|
The Naze
|
Spore
|
Cyatheaceae
|
|
| Cyathidites[42][44]
|
C. australis
|
The Naze
|
Spore
|
Cyatheaceae
|
|
| C. minor
|
Santa Marta Cove; The Naze
|
Spore
|
Cyatheaceae (Cyathea)
|
| Deltoidospora[42]
|
sp.
|
Santa Marta Cove
|
Spore
|
Fern affinity
|
|
| Densoisporites[44]
|
D. velatus
|
The Naze
|
Spore
|
Lycophyte affinity
|
|
| Dictyotosporites[44]
|
D. speciosus
|
The Naze
|
Spore
|
Fern affinity
|
|
| Echinosporis[42]
|
sp.
|
Santa Marta Cove
|
Spore
|
Fern affinity
|
|
| Gleicheniidites[44]
|
G. circinidites
|
Santa Marta Cove
|
Spore
|
Gleicheniaceae (aff. Gleichenia)
|
|
| G. senonicus
|
The Naze
|
Spore
|
Gleicheniaceae (aff. Gleichenia)
|
| Herkosporites[44]
|
sp.
|
The Naze
|
Spore
|
Fern affinity
|
|
| Ischyosporites[44]
|
I. volkheimeri
|
The Naze
|
Spore
|
Fern affinity
|
|
| cf. Klukisporites[44]
|
K. scaberis
|
The Naze
|
Spore
|
Lygodiaceae
|
|
| Laevigatosporites[44]
|
L. ovatus
|
Santa Marta Cove; The Naze
|
Spore
|
Polypodiaceae
|
|
| Leiotriletes[42]
|
sp.
|
Santa Marta Cove
|
Spore
|
Fern affinity
|
|
| Lycopodiumsporites[44]
|
L. eminulus
|
The Naze
|
Spore
|
Lycopodiaceae (Lycopodium)
|
|
| Millerocaulis[45]
|
M. santamartaensis
|
Santa Marta Cove
|
Fossil stem (hand specimens, slides)
|
Osmundaceae
|
|
| Peromonolites[44]
|
P. bowenii
|
The Naze
|
Spore
|
Fern affinity
|
|
| Perotrilites[44]
|
P. majus
|
The Naze
|
Spore
|
Fern affinity
|
|
| Polypodiisporites[44]
|
P. favus
|
The Naze
|
Spore
|
Polypodiaceae
|
|
| sp.
|
The Naze
|
Spore
|
Polypodiaceae
|
|
| Retitriletes[42]
|
R. austroclavatidites
|
Santa Marta Cove; The Naze
|
Spore
|
Lycopodiaceae (Lycopodium)
|
|
| Stereisporites[42][44]
|
S. antiquasporites
|
Santa Marta Cove; The Naze
|
Spore
|
Sphagnaceae (Moss)
|
|
| Stereisporites[44]
|
S. regium
|
The Naze
|
Spore
|
Moss affinity
|
|
| Todisporites[44]
|
T. major
|
The Naze
|
Spore
|
Fern affinity
|
|
| T. minor
|
The Naze
|
Spore
|
Fern affinity
|
|
| Trilites[42][44]
|
T. parvallatus
|
The Naze; Santa Marta Cove
|
Spore
|
Dicksoniaceae (Dicksonia)
|
|
| T. tuberculiformis
|
Santa Marta Cove
|
Spore
|
| Triporoletes[44]
|
T. radiatus
|
The Naze
|
Spore
|
Fern affinity
|
|
| Tuberculatosporites[44]
|
T. parvus
|
The Naze
|
Spore
|
Fern affinity
|
|
| Verrucosisporites[42]
|
sp. 2
|
Santa Marta Cove
|
Spore
|
Fern affinity
|
|
Gymnosperms (Conifers)
Angiosperms
See also
References
- ^ di Pasquo, M.; Martin, J.E. (2013). "Palynoassemblages Associated with a Theropod Dinosaur from the Snow Hill Island Formation (lower Maastrichtian) at the Naze, James Ross Island, Antarctica". Cretaceous Research. 45: 135–154. Bibcode:2013CrRes..45..135D. doi:10.1016/j.cretres.2013.07.008. hdl:11336/80121.
- ^ Ely & Case, 2019
- ^ O'Gorman, 2012, p.2
- ^ a b c d e f g h i j k l m n o p Reguero, Marcelo A.; Gasparini, Zulma; Olivero, Eduardo B.; Coria, Rodolfo A.; FernáNdez, Marta S.; O´Gorman, José P.; Gouiric-Cavalli, Soledad; Hospitaleche, Carolina Acosta; Bona, Paula; Iglesias, Ari; Gelfo, Javier N.; Raffi, MaríA E.; Moly, Juan José; Santillana, Sergio N.; CáRdenas, Magalí (2022). "Late Campanian-Early Maastrichtian Vertebrates From The James Ross Basin, West Antarctica: Updated Synthesis, Biostratigraphy, And Paleobiogeography". Anais da Academia Brasileira de Ciências. 94 (suppl 1) e20211142. doi:10.1590/0001-3765202220211142. ISSN 1678-2690. PMID 35674550.
- ^ O'Gorman, 2012, p.3
- ^ a b Videira-Santos, Roberto; Tobin, Thomas S.; Scheffler, Sandro Marcelo (2022-08-14). "New occurrences of caryophylliid and fungiacyathid scleractinian corals from the Santa Marta and Snow Hill Island formations (Upper Cretaceous, Antarctica)". Cretaceous Research. 140 105338. Bibcode:2022CrRes.14005338V. doi:10.1016/j.cretres.2022.105338. ISSN 0195-6671. S2CID 251576799.
- ^ a b c d Feldmann, Rodney M.; Tshudy, Dale M.; Thomson, Michael R. A. (1993). "Late Cretaceous and Paleocene Decapod Crustaceans from James Ross Basin, Antarctic Peninsula". Journal of Paleontology. 67 (S28): 1–41. Bibcode:1993JPal...67S...1F. doi:10.1017/S0022336000062077. ISSN 0022-3360.
- ^ a b c d e f g h i j k l Tobin, Thomas S.; Flannery, David; Sousa, Francis J. (2018-04-01). "Stratigraphy, sedimentology and paleontology of Upper Cretaceous deposits of Day Nunatak, Snow Hill Island, Antarctica". Cretaceous Research. 84: 407–419. Bibcode:2018CrRes..84..407T. doi:10.1016/j.cretres.2017.12.006. ISSN 0195-6671.
- ^ a b Hooker, J.J.; Milner, A.C.; Sequeira, S.E.K. (1991). "An ornithopod dinosaur from the Late Cretaceous of West Antarctica". Antarctic Science. 3 (3): 331–332. Bibcode:1991AntSc...3..331H. doi:10.1017/s0954102091000391. ISSN 0954-1020.
- ^ a b c d e f g h i Kriwet, J.; Lirio, J. M.; Nuñez, H. J.; Puceat, E.; Lécuyer, C. (2006-01-01), Francis, J. E.; Pirrie, D.; Crame, J. A. (eds.), "Late Cretaceous Antarctic fish diversity", Cretaceous–Tertiary High-Latitude Palaeoenvironments: James Ross Basin, Antarctica, vol. 258, Geological Society of London, p. 0, ISBN 978-1-86239-197-0, retrieved 2025-05-13
{{citation}}: CS1 maint: work parameter with ISBN (link)
- ^ Gallo, Valéria; de Figueiredo, Francisco J.; Brum, Arthur Souza; Soares, Marina Bento; Sayão, Juliana Manso; Pinheiro, André Eduardo Piacentini; Figueiredo, Rodrigo Giesta; Lopes, Ricardo Tadeu; de Araújo, Olga Maria Oliveira; Kellner, Alexander W. A. (2025-08-11). "First dercetid fish from the Upper Cretaceous of Antarctica". Scientific Reports. 15 (1): 26650. doi:10.1038/s41598-025-03570-y. ISSN 2045-2322.
- ^ a b c d e Otero, Rodrigo A.; Gutstein, Carolina Simon; Vargas, Alexander; Rubilar-Rogers, David; Yury-Yañez, Roberto; Bastías, Joaquin; Ramírez, Cristián (2014). "New Chondrichthyans from the Upper Cretaceous (Campanian–Maastrichtian) of Seymour and James Ross Islands, Antarctica". Journal of Paleontology. 88 (3): 411–420. doi:10.1666/13-041. ISSN 0022-3360.
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Bibliography
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Further reading
- D. Néraudeau, A. Crame, and M. Kooser. 2000. Upper Cretaceous echinoids from James Ross Basin, Antarctica. Géobios 33(4):455–466 doi:10.1016/S0016-6995(00)80079-0
- R. A. Otero, S. Soto-Acuna, A. O. Vargas, D. Rubilar Rogers, R. E. Yury Yanez and C. S. Gutstein. 2013. Additions to the diversity of elasmosaurid plesiosaurs from the Upper Cretaceous of Antarctica. Gondwana Research doi:10.1016/j.gr.2013.07.016