List of nesomyids

Nesomyidae is a family of mammals in the order Rodentia and part of the Myomorpha suborder. Members of this family are called nesomyids, and include Malagasy rodents, climbing mice, African rock mice, pouched rats, and tufted-tailed rats. They are found in sub-Saharan Africa, primarily in forests, savannas, shrublands, grasslands, and deserts, though some species can be found in rocky areas. They range in size from Monard's African climbing mouse, at 4 cm (2 in) plus a 5 cm (2 in) tail, to the southern giant pouched rat, at 41 cm (16 in) plus a 45 cm (18 in) tail. Nesomyids are generally omnivores, and eat seeds, fruit, nuts, roots, stems, and insects. Few nesomyids have population estimates, but five species—the white-tipped tufted-tailed rat, greater big-footed mouse, western nesomys, eastern voalavo, and Petter's tufted-tailed rat—are categorized as endangered, while the Mount Kahuzi climbing mouse and Malagasy giant rat are categorized as critically endangered.

The 67 extant species of Nesomyidae are divided into 21 genera, divided into 6 subfamilies. Cricetomyinae contains 8 species of hamster-rats and pouched rats in 3 genera, Delanymyinae contains a single species, Dendromurinae contains 26 species of climbing mice and fat mice in 6 genera, Mystromyinae contains a single species, Nesomyinae contains 27 species of short-tailed rats, tufted-tailed rats, and big-footed mice in 9 genera, and Petromyscinae contains 4 species of rock mice in a single genus. Several extinct prehistoric nesomyid species have been discovered, though due to ongoing research and discoveries, the exact number and categorization is not fixed.[1]

Conventions

IUCN Red List categories
Conservation status
 EX Extinct (0 species)
 EW Extinct in the wild (0 species)
 CR Critically endangered (2 species)
 EN Endangered (5 species)
 VU Vulnerable (5 species)
 NT Near threatened (0 species)
 LC Least concern (44 species)
Other categories
 DD Data deficient (8 species)
 NE Not evaluated (3 species)

The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the nesomyid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

Nesomyidae is a family consisting of 67 extant species in 21 genera. These genera are divided into six subfamilies: Cricetomyinae, containing 8 species of in 3 genera; Delanymyinae, containing a single species; Dendromurinae, containing 26 species in 6 genera; Mystromyinae, containing a single species; Nesomyinae, containing 27 species in 9 genera; and Petromyscinae, containing 4 species in a single genus. This does not include hybrid species or extinct prehistoric species.

Family Nesomyidae

Nesomyidae[2]

Nesomyids

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[3]

Subfamily Cricetomyinae

Genus Beamys Thomas, 1909 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Greater hamster-rat B. major
Dollman, 1914
Southeastern Africa Size: 13–18 cm (5–7 in) long, plus 12–15 cm (5–6 in) tail[4]

Habitat: Forest[5]

Diet: Seeds and fruit[5]
 NE 


Unknown

Lesser hamster-rat

B. hindei
Thomas, 1909
Southeastern Africa Size: 12–16 cm (5–6 in) long, plus 11–13 cm (4–5 in) tail[4]

Habitat: Forest[6]

Diet: Seeds and fruit[5]
 LC 


Unknown [6]

Genus Cricetomys Waterhouse, 1840 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Emin's pouched rat

C. emini
Wroughton, 1910
Central and western Africa Size: 30–35 cm (12–14 in) long, plus 32–43 cm (13–17 in) tail[4]

Habitat: Forest[7]

Diet: Omnivorous, including fruit and nuts as well as vegetables, insects, crabs, and snails[8]
 LC 


Unknown [7]

Gambian pouched rat

C. gambianus
Waterhouse, 1840
Central and western Africa Size: 28–39 cm (11–15 in) long, plus 25–40 cm (10–16 in) tail[4]

Habitat: Forest and savanna[9]

Diet: Omnivorous, including fruit and nuts as well as vegetables, insects, crabs, and snails[8]
 LC 


Unknown [9]

Kivu giant pouched rat C. kivuensis
Lönnberg, 1917
Central and western Africa Size: About 32 cm (13 in) long, plus about 34 cm (13 in) tail[4]

Habitat: Forest[7]

Diet: Omnivorous, including fruit and nuts as well as vegetables, insects, crabs, and snails[8]
 NE 


Unknown

Southern giant pouched rat

C. ansorgei
Thomas, 1904
Central and southern Africa
Size: 29–41 cm (11–16 in) long, plus 34–45 cm (13–18 in) tail[4]

Habitat: Forest and savanna[10]

Diet: Omnivorous, including fruit and nuts as well as vegetables, insects, crabs, and snails[8]
 LC 


Unknown [10]

Genus Saccostomus Peters, 1846 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Mearns's pouched mouse S. mearnsi
Heller, 1910
Eastern Africa Size: 10–15 cm (4–6 in) long, plus 3–8 cm (1–3 in) tail[4]

Habitat: Savanna, shrubland, and desert[11]

Diet: Seeds, berries, grains, and nuts, as well as insects[12]
 LC 


Unknown [11]

South African pouched mouse

S. campestris
Peters, 1846
Southern Africa Size: 9–18 cm (4–7 in) long, plus 2–6 cm (1–2 in) tail[4]

Habitat: Savanna, shrubland, grassland, and desert[13]

Diet: Seeds, berries, grains, and nuts, as well as insects[12]
 LC 


Unknown [13]

Subfamily Delanymyinae

Genus Delanymys Hayman, 1962 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Delany's mouse D. brooksi
Hayman, 1962
Central Africa
Size: 5–7 cm (2–3 in) long, plus 8–12 cm (3–5 in) tail[4]

Habitat: Shrubland and inland wetlands[14]

Diet: Seeds[5]
 VU 


Unknown [14]

Subfamily Dendromurinae

Genus Dendromus Smith, 1829 – fourteen species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Banana climbing mouse D. messorius
(Thomas, 1903)
Central and western Africa Size: 5–7 cm (2–3 in) long, plus 7–10 cm (3–4 in) tail[15]

Habitat: Grassland[16]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 LC 


Unknown [16]

Brants's climbing mouse

D. mesomelas
(Brants, 1827)
Southern Africa Size: 6–9 cm (2–4 in) long, plus 7–13 cm (3–5 in) tail[15]

Habitat: Forest, savanna, shrubland, and grassland[18]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 LC 


Unknown [18]

Cameroon climbing mouse D. oreas
Osgood, 1936
Cameroon Size: 6–8 cm (2–3 in) long, plus 8–11 cm (3–4 in) tail[15]

Habitat: Grassland[19]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 LC 


Unknown [19]

Chestnut climbing mouse

D. mystacalis
(Heuglin, 1863)
Eastern and southern Africa Size: 4–8 cm (2–3 in) long, plus 6–11 cm (2–4 in) tail[15]

Habitat: Savanna and grassland[20]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 LC 


Unknown [20]

Gray climbing mouse

D. melanotis
(Smith, 1834)
Sub-Saharan Africa Size: 5–8 cm (2–3 in) long, plus 5–10 cm (2–4 in) tail[15]

Habitat: Savanna, shrubland, grassland, and desert[21]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 LC 


Unknown [21]

Kivu climbing mouse D. nyasae
Thomas, 1916
Southern Africa Size: 6–8 cm (2–3 in) long, plus 8–11 cm (3–4 in) tail[15]

Habitat: Forest, savanna, shrubland, and grassland[18]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 NE 


Unknown

Lachaise's climbing mouse D. lachaisei
Denys & Aniskine, 2012
Western Africa
Size: 5–8 cm (2–3 in) long, plus 7–10 cm (3–4 in) tail[15]

Habitat: Forest and savanna[22]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 VU 


Unknown [22]

Lovat's climbing mouse

D. lovati
(de Winton, 1900)
Ethiopia Size: 5–10 cm (2–4 in) long, plus 5–9 cm (2–4 in) tail[15]

Habitat: Grassland[23]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 LC 


Unknown [23]

Monard's African climbing mouse D. leucostomus
Monard, 1933
Angola Size: 4–7 cm (2–3 in) long, plus 5–8 cm (2–3 in) tail[15]

Habitat: Savanna and grassland[17]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 NE 


Unknown

Montane African climbing mouse D. insignis
(Thomas, 1903)
East-central Africa Size: 7–10 cm (3–4 in) long, plus 8–12 cm (3–5 in) tail[15]

Habitat: Forest, shrubland, grassland, and inland wetlands[24]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 LC 


Unknown [24]

Mount Kahuzi climbing mouse D. kahuziensis
(Dieterlen, 1969)
Democratic Republic of the Congo Size: 7–9 cm (3–4 in) long, plus 12–14 cm (5–6 in) tail[15]

Habitat: Forest[25]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 CR 


Unknown [25]

Nyika climbing mouse D. nyikae
Wroughton, 1909
Southern Africa Size: 6–8 cm (2–3 in) long, plus 8–10 cm (3–4 in) tail[15]

Habitat: Forest, savanna, shrubland, and grassland[26]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 LC 


Unknown [26]

Rupp's African climbing mouse D. ruppi
Dieterlen, 2009
South Sudan Size: 6–9 cm (2–4 in) long, plus 9–12 cm (4–5 in) tail[15]

Habitat: Inland wetlands[27]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 DD 


Unknown [27]

Vernay's climbing mouse D. vernayi
Hill & Carter, 1937
Angola Size: 6–8 cm (2–3 in) long, plus 8–9 cm (3–4 in) tail[15]

Habitat: Savanna[28]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds[17]
 DD 


Unknown [28]

Genus Dendroprionomys Petter, 1966 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Velvet climbing mouse D. rousseloti
Petter, 1966
Republic of the Congo Size: About 8 cm (3 in) long, plus about 11 cm (4 in) tail[15]

Habitat: Unknown[29]

Diet: Insects[30]
 DD 


Unknown [29]

Genus Malacothrix Candolle, 1838 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Gerbil mouse M. typica
(Smith, 1834)
Southern Africa Size: 6–9 cm (2–4 in) long, plus 2–5 cm (1–2 in) tail[15]

Habitat: Desert, shrubland, and savanna[31]

Diet: Green vegetation[30]
 LC 


Unknown [31]

Genus Megadendromus Dieterlen & Rupp, 1978 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Nikolaus's mouse M. nikolausi
Dieterlen & Rupp, 1978
Ethiopia Size: 11–13 cm (4–5 in) long, plus 9–11 cm (4 in) tail[15]

Habitat: Shrubland[32]

Diet: Likely vegetation and insects[33]
 VU 


Unknown [32]

Genus Prionomys Dollman, 1910 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Dollman's tree mouse

P. batesi
Dollman, 1910
West-central Africa Size: 7–9 cm (3–4 in) long, plus 10–12 cm (4–5 in) tail[15]

Habitat: Forest[34]

Diet: Insects[30]
 DD 


Unknown [34]

Genus Steatomys Peters, 1846 – eight species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Bocage's African fat mouse S. bocagei
(Thomas, 1892)
Angola and Democratic Republic of the Congo Size: 9–13 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[15]

Habitat: Forest, savanna, and shrubland[35]

Diet: Seeds, grass bulbs, and insects[36]
 LC 


Unknown [35]

Dainty fat mouse S. cuppedius
Thomas & Hinton, 1920
Western Africa Size: 7–10 cm (3–4 in) long, plus 4–5 cm (2 in) tail[15]

Habitat: Shrubland and grassland[37]

Diet: Seeds, grass bulbs, and insects[36]
 LC 


Unknown [37]

Fat mouse

S. pratensis
Peters, 1846
Central and southern Africa Size: 8–11 cm (3–4 in) long, plus 4–6 cm (2 in) tail[15]

Habitat: Savanna and grassland[38]

Diet: Seeds, grass bulbs, and insects[36]
 LC 


Unknown [38]

Jackson's fat mouse S. jacksoni
Hayman, 1936
Ghana and Nigeria Size: About 12 cm (5 in) long, plus about 5 cm (2 in) tail[15]

Habitat: Unknown[39]

Diet: Seeds, grass bulbs, and insects[36]
 DD 


Unknown [39]

Krebs's fat mouse

S. krebsii
Peters, 1852
Southern Africa Size: 7–10 cm (3–4 in) long, plus 3–6 cm (1–2 in) tail[15]

Habitat: Shrubland and grassland[40]

Diet: Seeds, grass bulbs, and insects[36]
 LC 


Unknown [40]

Northwestern fat mouse S. caurinus
Thomas, 1912
Western Africa Size: 9–13 cm (4–5 in) long, plus 3–5 cm (1–2 in) tail[15]

Habitat: Shrubland[41]

Diet: Seeds, grass bulbs, and insects[36]
 LC 


Unknown [41]

Pousargues African fat mouse S. opimus
(Pousargues, 1894)
Central Africa Size: 11–13 cm (4–5 in) long, plus 5–7 cm (2–3 in) tail[15]

Habitat: Forest and savanna[42]

Diet: Seeds, grass bulbs, and insects[36]
 LC 


Unknown [42]

Tiny fat mouse S. parvus
Rhoads, 1896
Eastern and southern Africa Size: 5–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[15]

Habitat: Savanna, shrubland, and grassland[43]

Diet: Seeds, grass bulbs, and insects[36]
 LC 


Unknown [43]

Subfamily Mystromyinae

Genus Mystromys Wagner, 1841 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
White-tailed rat M. albicaudatus
(Smith, 1834)
Southern Africa Size: 11–20 cm (4–8 in) long, plus 4–9 cm (2–4 in) tail[4]

Habitat: Shrubland and grassland[44]

Diet: Seeds, vegetable matter, and insects[45]
 VU 


7,000–14,000 [44]

Subfamily Nesomyinae

Genus Brachytarsomys Günther, 1875 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Hairy-tailed antsangy B. villosa
Petter, 1962
Northern Madagascar
Size: 23–24 cm (9 in) long, plus 26–27 cm (10–11 in) tail[46]

Habitat: Forest[47]

Diet: Fruit[48]
 VU 


Unknown [47]

White-tailed antsangy

B. albicauda
Günther, 1875
Eastern Madagascar
Size: 22–23 cm (9 in) long, plus 22–24 cm (9 in) tail[46]

Habitat: Forest[49]

Diet: Fruit[48]
 LC 


Unknown [49]

Genus Brachyuromys Forsyth Major, 1896 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Betsileo short-tailed rat B. betsileoensis
(Bartlett, 1880)
Eastern Madagascar
Size: 14–19 cm (6–7 in) long, plus 7–10 cm (3–4 in) tail[4]

Habitat: Forest, shrubland, grassland, and inland wetlands[50]

Diet: Likely vegetation and insects[33]
 LC 


Unknown [50]

Gregarious short-tailed rat B. ramirohitra
Forsyth Major, 1896
Eastern Madagascar
Size: 14–17 cm (6–7 in) long, plus 8–11 cm (3–4 in) tail[4]

Habitat: Forest[51]

Diet: Likely vegetation and insects[33]
 LC 


Unknown [51]

Genus Eliurus A. Milne-Edwards, 1885 – twelve species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Ankarana Special Reserve tufted-tailed rat E. carletoni
Goodman, Raheriarisena, & Jansa, 2009
Northern Madagascar Size: 14–15 cm (6 in) long, plus 16–19 cm (6–7 in) tail[46]

Habitat: Forest[52]

Diet: Likely vegetation and insects[33]
 LC 


Unknown [52]

Daniel's tufted-tailed rat E. danieli
Goodman & Carleton, 2007
Southern Madagascar
Size: 15–16 cm (6 in) long, plus 17–20 cm (7–8 in) tail[46]

Habitat: Shrubland[53]

Diet: Likely vegetation and insects[33]
 LC 


Unknown [53]

Dormouse tufted-tailed rat

E. myoxinus
A. Milne-Edwards, 1855
Northern, western, and southern Madagascar
Size: 11–14 cm (4–6 in) long, plus 12–17 cm (5–7 in) tail[46]

Habitat: Forest[54]

Diet: Likely vegetation and insects[33]
 LC 


Unknown [54]

Ellerman's tufted-tailed rat E. ellermani
Carleton, 1994
Eastern Madagascar
Size: About 15 cm (6 in) long, plus about 18 cm (7 in) tail[46]

Habitat: Forest[55]

Diet: Likely vegetation and insects[33]
 DD 


Unknown [55]

Grandidier's tufted-tailed rat E. grandidieri
Goodman & Carleton, 1998
Northeastern Madagascar
Size: 11–17 cm (4–7 in) long, plus 14–18 cm (6–7 in) tail[46]

Habitat: Forest[56]

Diet: Likely vegetation and insects[33]
 LC 


Unknown [56]

Lesser tufted-tailed rat E. minor
Forsyth Major, 1896
Northern and eastern Madagascar
Size: 10–13 cm (4–5 in) long, plus 11–14 cm (4–6 in) tail[46]

Habitat: Forest[57]

Diet: Likely vegetation and insects[33]
 LC 


Unknown [57]

Major's tufted-tailed rat E. majori
Thomas, 1895
Eastern Madagascar
Size: 13–17 cm (5–7 in) long, plus 15–20 cm (6–8 in) tail[46]

Habitat: Forest[58]

Diet: Likely vegetation and insects[33]
 LC 


Unknown [58]

Petter's tufted-tailed rat E. petteri
Carleton, 1994
Eastern Madagascar Size: About 13 cm (5 in) long, plus about 19 cm (7 in) tail[46]

Habitat: Forest[59]

Diet: Likely vegetation and insects[33]
 EN 


Unknown [59]

Tanala tufted-tailed rat E. tanala
Forsyth Major, 1896
Eastern Madagascar
Size: 14–16 cm (6 in) long, plus 15–20 cm (6–8 in) tail[46]

Habitat: Forest[60]

Diet: Likely vegetation and insects[33]
 LC 


Unknown [60]

Tsingy tufted-tailed rat E. antsingy
Goodman, Rakotondravony, & Carleton, 2001
Northern Madagascar
Size: 14–16 cm (6 in) long, plus 15–20 cm (6–8 in) tail[46]

Habitat: Forest[61]

Diet: Likely vegetation and insects[33]
 DD 


Unknown [61]

Webb's tufted-tailed rat E. webbi
Ellerman, 1949
Eastern Madagascar
Size: 14–16 cm (6 in) long, plus 16–19 cm (6–7 in) tail[46]

Habitat: Forest[62]

Diet: Likely vegetation and insects[33]
 LC 


Unknown [62]

White-tipped tufted-tailed rat E. penicillatus
Thomas, 1908
East-central Madagascar
Size: About 15 cm (6 in) long, plus about 17 cm (7 in) tail[46]

Habitat: Forest[63]

Diet: Likely vegetation and insects[33]
 EN 


Unknown [63]

Genus Gymnuromys Forsyth Major, 1896 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Voalavoanala G. roberti
Forsyth Major, 1896
Eastern Madagascar
Size: 15–18 cm (6–7 in) long, plus 14–20 cm (6–8 in) tail[46]

Habitat: Forest[64]

Diet: Likely vegetation and insects[33]
 LC 


Unknown [64]

Genus Hypogeomys Grandidier, 1869 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Malagasy giant rat

H. antimena
Grandidier, 1869
Western Madagascar
Size: 30–34 cm (12–13 in) long, plus 21–24 cm (8–9 in) tail[46]

Habitat: Forest[65]

Diet: Fruit[66]
 CR 


5,000 [65]

Genus Macrotarsomys A. Milne-Edwards & Grandidier, 1898 – three species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Bastard big-footed mouse

M. bastardi
A. Milne-Edwards & Grandidier, 1898
Western Madagascar
Size: 8–11 cm (3–4 in) long, plus 12–15 cm (5–6 in) tail[46]

Habitat: Forest and shrubland[67]

Diet: Berries, fruit, seeds, roots, and stems[48]
 LC 


Unknown [67]

Greater big-footed mouse M. ingens
(Petter, 1959)
Northern Madagascar
Size: 11–15 cm (4–6 in) long, plus 18–24 cm (7–9 in) tail[46]

Habitat: Forest[68]

Diet: Berries, fruit, seeds, roots, and stems[48]
 EN 


Unknown [68]

Petter's big-footed mouse M. petteri
Goodman & Soarimalala, 2005
Southern Madagascar
Size: About 16 cm (6 in) long, plus about 24 cm (9 in) tail[46]

Habitat: Forest[69]

Diet: Berries, fruit, seeds, roots, and stems[48]
 DD 


Unknown [69]

Genus Monticolomys Goodman & Carleton, 1996 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Malagasy mountain mouse M. koopmani
Goodman & Carleton, 1996
Northern, central, and southern Madagascar
Size: 9–11 cm (4 in) long, plus 13–15 cm (5–6 in) tail[46]

Habitat: Forest[70]

Diet: Likely vegetation and insects[33]
 LC 


Unknown [70]

Genus Nesomys Peters, 1870 – three species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Island mouse

N. rufus
Peters, 1870
Eastern Madagascar
Size: 17–20 cm (7–8 in) long, plus 16–18 cm (6–7 in) tail[4]

Habitat: Forest[71]

Diet: Fruit and nuts[48]
 LC 


Unknown [71]

Western nesomys

N. lambertoni
Grandidier, 1928
Western Madagascar Size: 18–20 cm (7–8 in) long, plus 18–20 cm (7–8 in) tail[4]

Habitat: Forest[72]

Diet: Fruit and nuts[48]
 EN 


Unknown [72]

White-bellied nesomys

N. audeberti
Jentink, 1879
Eastern Madagascar Size: 19–20 cm (7–8 in) long, plus 16–18 cm (6–7 in) tail[4]

Habitat: Forest[73]

Diet: Fruit and nuts[48]
 LC 


Unknown [73]

Genus Voalavo Goodman & Carleton, 1998 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Eastern voalavo V. antsahabensis
Goodman, Rakotondravony, Randriamanantsoa, & Rakotomalala-Razanahoera, 2005
Central Madagascar (in green)
Size: 8–10 cm (3–4 in) long, plus 10–13 cm (4–5 in) tail[46]

Habitat: Forest[74]

Diet: Likely vegetation and insects[33]
 EN 


Unknown [74]

Northern voalavo V. gymnocaudus
Goodman & Carleton, 1998
Northern Madagascar (in red)
Size: 8–9 cm (3–4 in) long, plus 12–13 cm (5 in) tail[46]

Habitat: Forest[75]

Diet: Likely vegetation and insects[33]
 LC 


Unknown [75]

Subfamily Petromyscinae

Genus Petromyscus Thomas, 1926 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Barbour's rock mouse P. barbouri
Shortridge, 1938
South Africa Size: 7–8 cm (3 in) long, plus 7–9 cm (3–4 in) tail[4]

Habitat: Shrubland and rocky areas[76]

Diet: Omnivorous[5]
 LC 


Unknown [76]

Brukkaros pygmy rock mouse P. monticularis
(Thomas & Hinton, 1925)
Namibia and South Africa Size: 6–10 cm (2–4 in) long, plus 6–9 cm (2–4 in) tail[4]

Habitat: Shrubland and rocky areas[77]

Diet: Omnivorous[5]
 LC 


Unknown [77]

Pygmy rock mouse P. collinus
(Hinton, 1925)
Southwestern Africa Size: 7–10 cm (3–4 in) long, plus 7–13 cm (3–5 in) tail[4]

Habitat: Shrubland and rocky areas[78]

Diet: Omnivorous[5]
 LC 


Unknown [78]

Shortridge's rock mouse P. shortridgei
Thomas, 1926
Angola and Namibia Size: 6–10 cm (2–4 in) long, plus 7–11 cm (3–4 in) tail[4]

Habitat: Shrubland and rocky areas[79]

Diet: Omnivorous[5]
 LC 


Unknown [79]

References

  1. ^ "Family Nesomyidae (mouse)". Paleobiology Database. University of Wisconsin–Madison. Archived from the original on November 26, 2024. Retrieved September 18, 2025.
  2. ^ Steppan, S. J.; Schenk, J. J. (2017). "Muroid rodent phylogenetics: 900-species tree reveals increasing diversification rates". PLOS One. 12 (8) e0183070. doi:10.1371/journal.pone.0183070. PMC 5559066.
  3. ^ Wilson; Reeder, pp. 930–954
  4. ^ a b c d e f g h i j k l m n o p q r s Chernasky; Motis; Burgin, pp. 216–217
  5. ^ a b c d e f g h Nowak, pp. 1492–1493
  6. ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Beamys hindei". IUCN Red List of Threatened Species. 2016 e.T2676A115063057. doi:10.2305/IUCN.UK.2016-3.RLTS.T2676A22238241.en.
  7. ^ a b c Cassola, F. (2017) [errata version of 2016 assessment]. "Cricetomys emini". IUCN Red List of Threatened Species. 2016 e.T5521A115072329. doi:10.2305/IUCN.UK.2016-3.RLTS.T5521A22236579.en.
  8. ^ a b c d Nowak, p. 1496
  9. ^ a b Kennerley, R. (2023) [amended version of 2019 assessment]. "Cricetomys gambianus". IUCN Red List of Threatened Species. 2023 e.T112169507A243173564. doi:10.2305/IUCN.UK.2023-1.RLTS.T112169507A243173564.en.
  10. ^ a b Dando, T.; Kennerley, R. (2023) [amended version of 2019 assessment]. "Cricetomys ansorgei". IUCN Red List of Threatened Species. 2023 e.T112256552A243175763. doi:10.2305/IUCN.UK.2023-1.RLTS.T112256552A243175763.en.
  11. ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Saccostomus mearnsi". IUCN Red List of Threatened Species. 2016 e.T19809A115153561. doi:10.2305/IUCN.UK.2016-3.RLTS.T19809A22234073.en.
  12. ^ a b Nowak, p. 1495
  13. ^ a b Cassola, F.; Child, M. F. (2017) [errata version of 2016 assessment]. "Saccostomus campestris". IUCN Red List of Threatened Species. 2016 e.T19808A115153363. doi:10.2305/IUCN.UK.2016-3.RLTS.T19808A22233928.en.
  14. ^ a b Kennerley, R.; Kerbis Peterhans, J. (2020) [errata version of 2016 assessment]. "Delanymys brooksi". IUCN Red List of Threatened Species. 2016 e.T6313A184098557. doi:10.2305/IUCN.UK.2016-3.RLTS.T6313A184098557.en.
  15. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z Chernasky; Motis; Burgin, pp. 218–219
  16. ^ a b Schlitter, D.; Dieterlen, F. (2017) [errata version of 2016 assessment]. "Dendromus messorius". IUCN Red List of Threatened Species. 2016 e.T6445A115081283. doi:10.2305/IUCN.UK.2016-3.RLTS.T6445A22235137.en.
  17. ^ a b c d e f g h i j k l m n o Nowak, p. 1487
  18. ^ a b c Child, M. F.; Monadjem, A. (2016). "Dendromus mesomelas". IUCN Red List of Threatened Species. 2016 e.T6444A22235226. doi:10.2305/IUCN.UK.2016-3.RLTS.T6444A22235226.en.
  19. ^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Dendromus oreas". IUCN Red List of Threatened Species. 2016 e.T6448A115081767. doi:10.2305/IUCN.UK.2016-3.RLTS.T6448A22234774.en.
  20. ^ a b Child, M. F.; Monadjem, A. (2017) [errata version of 2016 assessment]. "Dendromus mystacalis". IUCN Red List of Threatened Species. 2016 e.T6446A115081411. doi:10.2305/IUCN.UK.2016-3.RLTS.T6446A22234993.en.
  21. ^ a b Child, M. F.; Monadjem, A. (2016). "Dendromus melanotis". IUCN Red List of Threatened Species. 2016 e.T6443A22235350. doi:10.2305/IUCN.UK.2016-3.RLTS.T6443A22235350.en.
  22. ^ a b Denys, C.; Kennerley, R. (2024). "Dendromus lachaisei". IUCN Red List of Threatened Species. 2024 e.T45958985A258446545. doi:10.2305/IUCN.UK.2024-2.RLTS.T45958985A258446545.en.
  23. ^ a b Lavrenchenko, L. (2016). "Dendromus lovati". IUCN Red List of Threatened Species. 2016 e.T6442A22235513. doi:10.2305/IUCN.UK.2016-2.RLTS.T6442A22235513.en.
  24. ^ a b Cassola, F. (2016). "Dendromus insignis". IUCN Red List of Threatened Species. 2016 e.T6440A22234456. doi:10.2305/IUCN.UK.2016-2.RLTS.T6440A22234456.en.
  25. ^ a b Dando, T.; Demos, T. (2021). "Dendromus kahuziensis". IUCN Red List of Threatened Species. 2021 e.T6439A22234587. doi:10.2305/IUCN.UK.2021-2.RLTS.T6439A22234587.en.
  26. ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Dendromus nyikae". IUCN Red List of Threatened Species. 2016 e.T6447A115081603. doi:10.2305/IUCN.UK.2016-3.RLTS.T6447A22234880.en.
  27. ^ a b Denys, C.; Kennerley, R. (2024). "Dendromus ruppi". IUCN Red List of Threatened Species. 2024 e.T45959000A258446746. doi:10.2305/IUCN.UK.2024-2.RLTS.T45959000A258446746.en.
  28. ^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Dendromus vernayi". IUCN Red List of Threatened Species. 2016 e.T6449A115081904. doi:10.2305/IUCN.UK.2016-3.RLTS.T6449A22234694.en.
  29. ^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Dendroprionomys rousseloti". IUCN Red List of Threatened Species. 2016 e.T6483A115082012. doi:10.2305/IUCN.UK.2016-3.RLTS.T6483A22234171.en.
  30. ^ a b c Nowak, pp. 1488–1489
  31. ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Malacothrix typica". IUCN Red List of Threatened Species. 2016 e.T12712A115105728. doi:10.2305/IUCN.UK.2016-3.RLTS.T12712A22238026.en.
  32. ^ a b Gerrie, R.; Kennerley, R. (2019). "Megadendromus nikolausi". IUCN Red List of Threatened Species. 2019 e.T12937A22237949. doi:10.2305/IUCN.UK.2019-1.RLTS.T12937A22237949.en.
  33. ^ a b c d e f g h i j k l m n o p q r s Nowak, p. 1346
  34. ^ a b Gerrie, R.; Kennerley, R. (2016). "Prionomys batesi". IUCN Red List of Threatened Species. 2016 e.T18164A22236213. doi:10.2305/IUCN.UK.2016-3.RLTS.T18164A22236213.en.
  35. ^ a b Hoffmann, M.; Cox, N. A. (2017). "Steatomys bocagei". IUCN Red List of Threatened Species. 2017 e.T136625A22233115. doi:10.2305/IUCN.UK.2017-2.RLTS.T136625A22233115.en.
  36. ^ a b c d e f g h Nowak, p. 1490
  37. ^ a b Schlitter, D.; Coetzee, N. (2017). "Steatomys cuppedius". IUCN Red List of Threatened Species. 2017 e.T20719A22233546. doi:10.2305/IUCN.UK.2017-2.RLTS.T20719A22233546.en.
  38. ^ a b Child, M. F.; Monadjem, A. (2017) [errata version of 2016 assessment]. "Steatomys pratensis". IUCN Red List of Threatened Species. 2016 e.T20722A115159593. doi:10.2305/IUCN.UK.2016-3.RLTS.T20722A22233313.en.
  39. ^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Steatomys jacksoni". IUCN Red List of Threatened Species. 2016 e.T20717A115159338. doi:10.2305/IUCN.UK.2016-3.RLTS.T20717A22233758.en.
  40. ^ a b Monadjem, A.; Schlitter, D. (2017) [errata version of 2016 assessment]. "Steatomys krebsii". IUCN Red List of Threatened Species. 2016 e.T20720A115159464. doi:10.2305/IUCN.UK.2016-3.RLTS.T20720A22233457.en.
  41. ^ a b Schlitter, D.; Coetzee, N. (2017). "Steatomys caurinus". IUCN Red List of Threatened Species. 2017 e.T20718A22233646. doi:10.2305/IUCN.UK.2017-2.RLTS.T20718A22233646.en.
  42. ^ a b Hoffmann, M.; Cox, N. A. (2017). "Steatomys opimus". IUCN Red List of Threatened Species. 2017 e.T136513A22233848. doi:10.2305/IUCN.UK.2017-2.RLTS.T136513A22233848.en.
  43. ^ a b Schlitter, D.; Monadjem, A. (2017). "Steatomys parvus". IUCN Red List of Threatened Species. 2017 e.T20721A22233188. doi:10.2305/IUCN.UK.2017-2.RLTS.T20721A22233188.en.
  44. ^ a b Avenant, N. L.; Wilson, B.; Power, J.; Palmer, G.; Child, M. F. (2019). "Mystromys albicaudatus". IUCN Red List of Threatened Species. 2019 e.T14262A22237378. doi:10.2305/IUCN.UK.2019-1.RLTS.T14262A22237378.en.
  45. ^ Nowak, p. 1425
  46. ^ a b c d e f g h i j k l m n o p q r s t u v Chernasky; Motis; Burgin, pp. 214–215
  47. ^ a b Kennerley, R.; Goodman, S. (2016). "Brachytarsomys villosa". IUCN Red List of Threatened Species. 2016 e.T136399A22237177. doi:10.2305/IUCN.UK.2016-3.RLTS.T136399A22237177.en.
  48. ^ a b c d e f g h Nowak, pp. 1434–1435
  49. ^ a b Kennerley, R. (2016). "Brachytarsomys albicauda". IUCN Red List of Threatened Species. 2016 e.T2991A22237052. doi:10.2305/IUCN.UK.2016-2.RLTS.T2991A22237052.en.
  50. ^ a b Kennerley, R. (2016). "Brachyuromys betsileoensis". IUCN Red List of Threatened Species. 2016 e.T2997A22234256. doi:10.2305/IUCN.UK.2016-2.RLTS.T2997A22234256.en.
  51. ^ a b Kennerley, R. (2016). "Brachyuromys ramirohitra". IUCN Red List of Threatened Species. 2016 e.T2998A22234363. doi:10.2305/IUCN.UK.2016-2.RLTS.T2998A22234363.en.
  52. ^ a b Kennerley, R. (2016). "Eliurus carletoni". IUCN Red List of Threatened Species. 2016 e.T48266951A48266956. doi:10.2305/IUCN.UK.2016-3.RLTS.T48266951A48266956.en.
  53. ^ a b Hoffmann, M.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Eliurus danieli". IUCN Red List of Threatened Species. 2016 e.T136251A115205056. doi:10.2305/IUCN.UK.2016-3.RLTS.T136251A22238907.en.
  54. ^ a b Goodman, S. (2016). "Eliurus myoxinus". IUCN Red List of Threatened Species. 2016 e.T7623A22238673. doi:10.2305/IUCN.UK.2016-2.RLTS.T7623A22238673.en.
  55. ^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Eliurus ellermani". IUCN Red List of Threatened Species. 2016 e.T136325A115205656. doi:10.2305/IUCN.UK.2016-3.RLTS.T136325A22238824.en.
  56. ^ a b Kennerley, R. (2016). "Eliurus grandidieri". IUCN Red List of Threatened Species. 2016 e.T29460A22238512. doi:10.2305/IUCN.UK.2016-3.RLTS.T29460A22238512.en.
  57. ^ a b Kennerley, R. (2016). "Eliurus minor". IUCN Red List of Threatened Species. 2016 e.T7621A22238349. doi:10.2305/IUCN.UK.2016-2.RLTS.T7621A22238349.en.
  58. ^ a b Kennerley, R. (2016). "Eliurus majori". IUCN Red List of Threatened Species. 2016 e.T7622A22238442. doi:10.2305/IUCN.UK.2016-2.RLTS.T7622A22238442.en.
  59. ^ a b Kennerley, R.; Goodman, S. (2016). "Eliurus petteri". IUCN Red List of Threatened Species. 2016 e.T136316A22238985. doi:10.2305/IUCN.UK.2016-3.RLTS.T136316A22238985.en.
  60. ^ a b Kennerley, R. (2016). "Eliurus tanala". IUCN Red List of Threatened Species. 2016 e.T7625A22239073. doi:10.2305/IUCN.UK.2016-2.RLTS.T7625A22239073.en.
  61. ^ a b Kennerley, R. (2016). "Eliurus antsingy". IUCN Red List of Threatened Species. 2016 e.T48267436A22238593. doi:10.2305/IUCN.UK.2016-3.RLTS.T48267436A22238593.en.
  62. ^ a b Kennerley, R. (2016). "Eliurus webbi". IUCN Red List of Threatened Species. 2016 e.T7626A22239153. doi:10.2305/IUCN.UK.2016-2.RLTS.T7626A22239153.en.
  63. ^ a b Kennerley, R. (2016). "Eliurus penicillatus". IUCN Red List of Threatened Species. 2016 e.T7624A22238746. doi:10.2305/IUCN.UK.2016-3.RLTS.T7624A22238746.en.
  64. ^ a b Kennerley, R. (2016). "Gymnuromys roberti". IUCN Red List of Threatened Species. 2016 e.T9581A22237297. doi:10.2305/IUCN.UK.2016-2.RLTS.T9581A22237297.en.
  65. ^ a b Wilmet, L.; Volahy, A.; Hudson, M. A. (2022). "Hypogeomys antimena". IUCN Red List of Threatened Species. 2022 e.T10714A216087357. doi:10.2305/IUCN.UK.2022-1.RLTS.T10714A216087357.en.
  66. ^ Nowak, p. 1437
  67. ^ a b Kennerley, R. (2016). "Macrotarsomys bastardi". IUCN Red List of Threatened Species. 2016 e.T12647A22236959. doi:10.2305/IUCN.UK.2016-2.RLTS.T12647A22236959.en.
  68. ^ a b Dollar, L.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Macrotarsomys ingens". IUCN Red List of Threatened Species. 2016 e.T12646A115105272. doi:10.2305/IUCN.UK.2016-3.RLTS.T12646A22236876.en.
  69. ^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Macrotarsomys petteri". IUCN Red List of Threatened Species. 2016 e.T136510A115209149. doi:10.2305/IUCN.UK.2016-3.RLTS.T136510A22236785.en.
  70. ^ a b Kennerley, R. (2016). "Monticolomys koopmani". IUCN Red List of Threatened Species. 2016 e.T136539A22233042. doi:10.2305/IUCN.UK.2016-3.RLTS.T136539A22233042.en.
  71. ^ a b Kennerley, R. (2016). "Nesomys rufus". IUCN Red List of Threatened Species. 2016 e.T14671A22237658. doi:10.2305/IUCN.UK.2016-2.RLTS.T14671A22237658.en.
  72. ^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Nesomys lambertoni". IUCN Red List of Threatened Species. 2016 e.T136330A115206046. doi:10.2305/IUCN.UK.2016-3.RLTS.T136330A22237750.en.
  73. ^ a b Kennerley, R. (2016). "Nesomys audeberti". IUCN Red List of Threatened Species. 2016 e.T136475A22237564. doi:10.2305/IUCN.UK.2016-2.RLTS.T136475A22237564.en.
  74. ^ a b Kennerley, R. (2019). "Voalavo antsahabensis". IUCN Red List of Threatened Species. 2019 e.T165925A22236108. doi:10.2305/IUCN.UK.2019-1.RLTS.T165925A22236108.en.
  75. ^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Voalavo gymnocaudus". IUCN Red List of Threatened Species. 2016 e.T29461A115169008. doi:10.2305/IUCN.UK.2016-3.RLTS.T29461A22236010.en.
  76. ^ a b Schlitter, D. (2016). "Petromyscus barbouri". IUCN Red List of Threatened Species. 2016 e.T16777A22235766. doi:10.2305/IUCN.UK.2016-2.RLTS.T16777A22235766.en.
  77. ^ a b Child, M. F. (2017) [errata version of 2016 assessment]. "Petromyscus monticularis". IUCN Red List of Threatened Species. 2016 e.T16779A115139351. doi:10.2305/IUCN.UK.2016-3.RLTS.T16779A22235937.en.
  78. ^ a b Child, M. F. (2017) [errata version of 2016 assessment]. "Petromyscus collinus". IUCN Red List of Threatened Species. 2016 e.T16778A115139225. doi:10.2305/IUCN.UK.2016-3.RLTS.T16778A22235686.en.
  79. ^ a b Schlitter, D. (2016). "Petromyscus shortridgei". IUCN Red List of Threatened Species. 2016 e.T16780A22235864. doi:10.2305/IUCN.UK.2016-2.RLTS.T16780A22235864.en.

Sources