Hypselostoma

Hypselostoma
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Family:	Hypselostomatidae
Genus:	Hypselostoma; Benson, 1856
Synonyms
	Anthroapiculus Panha & J. B. Burch, 2002 (incorrect subsequent spelling); Antroapiculus Panha & J. B. Burch, 2002 superseded combination; Boysidia (Dasypupa) F. G. Thompson & Dance, 1983 junior subjective synonym; Gyliauchen Pilsbry, 1917 (invalid: junior homonym of Gyliauchen Nicoll, 1915 [Plathelminthes]; Gyliotrachela is a replacement name); Gyliotrachela Tomlin, 1930 junior subjective synonym; Tanystoma W. H. Benson, 1856 (Invalid: junior homonym of Tanystoma Motschoulsky, 1845 [Coleoptera]; Hypselostoma is a replacement name);

Hypselostoma is a genus of very small air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Hypselostomatidae, the whorl snails.

Some species have the aperture of the shell on a trumpet-shaped extension.^[2]

Species

Species within the genus Hypselostoma include:^[3]

Hypselostoma adela F. G. Thompson & Upatham, 1997
Hypselostoma aenigma Gojšina, Grego & Páll-Gergely, 2025
Hypselostoma annamiticum Möllendorff, 1900
Hypselostoma aquila Gojšina, Hunyadi & Páll-Gergely, 2025
Hypselostoma aunglini (Tongkerd & Panha, 2024)
Hypselostoma australe Odhner, 1917
Hypselostoma benetuitum Vermeulen, Luu, Theary & Anker, 2019^[4]
Hypselostoma bensonianum Blanford, 1863
Hypselostoma bubalus Gojšina, Hunyadi & Páll-Gergely, 2025
Hypselostoma burchi Panha, 1997
Hypselostoma cambodjense Van Benthem Jutting, 1962
Hypselostoma chatnareeae (Panha & J. B. Burch, 2003)
Hypselostoma chaunosalpinx (Vermeulen, Luu, Theary & Anker, 2019)
Hypselostoma chedi Panha, 1998
Hypselostoma circumcarinatum Gojšina, Auffenberg & Páll-Gergely, 2025
Hypselostoma coriaceum Gojšina & Páll-Gergely, 2025
Hypselostoma concreta van Benthem Jutting, 1949
Hypselostoma crossei (Morlet, 1886)
Hypselostoma cucumense Panha, 1997
Hypselostoma cultura Tanmuangpak & Dumrongrojwattana, 2022
Hypselostoma depressispira van Benthem Jutting, 1949
Hypselostoma diarmaidi Panha & J. B. Burch, 2003
Hypselostoma dilatatum van Benthem Jutting, 1962
Hypselostoma discobasis Vermeulen, Luu, Theary & Anker, 2019
Hypselostoma edentatum Panha & J. B. Burch, 1999
Hypselostoma edentulum Moellendorff, 1894
Hypselostoma elephas van Benthem Jutting, 1950
Hypselostoma emergens van Benthem Jutting, 1950
Hypselostoma erawan Panha & J. B. Burch, 2003
Hypselostoma erawan Panha & J. B. Burch, 2002
Hypselostoma everetti (E. A. Smith, 1896)
Hypselostoma fortunatum Gojšina, Hunyadi & Páll-Gergely, 2025
Hypselostoma frequens (van Benthem Jutting, 1950)
Hypselostoma fruhstorferi (Möllendorff, 1897)
Hypselostoma fungus Gojšina, Hunyadi & Páll-Gergely, 2025
Hypselostoma geckophilum Gojšina, Hunyadi & Páll-Gergely, 2025
Hypselostoma holimanae F. G. Thompson & Lee, 1988
Hypselostoma hungerfordianum (Möllendorff, 1891)
Hypselostoma insularum Pilsbry, 1908
Hypselostoma iunior Gojšina & Páll-Gergely, 2025
Hypselostoma kelantanense (Sykes, 1902)
Hypselostoma khaochakan Panha & J. B. Burch, 2003
Hypselostoma khaochongpran Panha & J. B. Burch, 2002^[5]
Hypselostoma khaochongensis Panha, 1997
Hypselostoma khaowongense (Panha, 1997)
Hypselostoma khaowongkot Panha & J. B. Burch, 2004
Hypselostoma khmerianum (Sutcharit & Panha, 2023)
Hypselostoma kohrin Panha & J. B. Burch, 2003
Hypselostoma latispira F. G. Thompson & Auffenberg, 1984
Hypselostoma lipaei Gojšina, Schilthuizen & Páll-Gergely, 2023
Hypselostoma loei Panha & Prateespasen, 2005
Hypselostoma luctans van Benthem Jutting, 1950
Hypselostoma luzonicum Moellendorff, 1888
Hypselostoma modestum van Benthem Jutting, 1950
Hypselostoma muaklekense (Panha & J. B. Burch, 2002)
Hypselostoma muangon Panha & J. B. Burch, 2004
Hypselostoma muaklekense (Panha & J. B. Burch, 2002)
Hypselostoma paini (F. G. Thompson & Dance, 1983)
Hypselostoma panhai J. B. Burch & Tongkerd, 2002
Hypselostoma pattalungense Panha & J. B. Burch, 2004^[6]
Hypselostoma pendulum (Panha & J. B. Burch, 2002)
Hypselostoma phoca Tongkerd & Panha, 2013
Hypselostoma phupaman (Panha & J. B. Burch, 2002)
Hypselostoma piconis van Benthem-Jutting, 1949
Hypselostoma platybasis Gojšina, Hunyadi & Páll-Gergely, 2025
Hypselostoma plesiolopa Inkhavilay & Panha, 2016
Hypselostoma polyodon Moellendorff, 1896
Hypselostoma pongrati Tanmuangpak & S. Tumpeesuwan, 2025
Hypselostoma populare Gojšina, Hunyadi & Páll-Gergely, 2025
Hypselostoma procerum (F. G. Thompson & Dance, 1983)
Hypselostoma pusillum Moellendorff, 1894
Hypselostoma quadrasi Moellendorff, 1896
Hypselostoma roebeleni Moellendorff, 1894
Hypselostoma rupestre van Benthem Jutting, 1962
Hypselostoma salpinx van Benthem Jutting, 1961
Hypselostoma saraburiensis Panha & J. B. Burch, 2003
Hypselostoma saxicola van Benthem Jutting, 1960
Hypselostoma sculpturatum Gojšina, Hunyadi & Páll-Gergely, 2025
Hypselostoma serpa (van Benthem Jutting, 1950)
Hypselostoma sibuyanicum Moellendorff, 1896
Hypselostoma sichang (Panha & J. B. Burch, 2002)
Hypselostoma sichomphuense Tanmuangpak & S. Tumpeesuwan, 2025
Hypselostoma sigma Gojšina, Schilthuizen & Páll-Gergely, 2023
Hypselostoma similare Gojšina, Hunyadi & Páll-Gergely, 2025
Hypselostoma smokon Panha & J. B. Burch, 2004
Hypselostoma sorormajor Gojšina, Hunyadi & Páll-Gergely, 2025
Hypselostoma sororminor Gojšina, Hunyadi & Páll-Gergely, 2025
Hypselostoma srakeoensis (Panha & J. B. Burch, 2004)
Hypselostoma srirachaensis Panha & J. B. Burch, 2004
Hypselostoma surakiti Panha & J. B. Burch, 2003
Hypselostoma taehwani Panha & J. B. Burch, 2002
Hypselostoma tarutao (Panha & J. B. Burch, 2002)
Hypselostoma terae Tomlin, 1939
Hypselostoma tertiusfrater Gojšina & Páll-Gergely, 2025
Hypselostoma torta Gojšina, Auffenberg & Páll-Gergely, 2025
Hypselostoma torticollis van Benthem Jutting, 1962
Hypselostoma transitans (Möllendorff, 1894)
Hypselostoma tridentatum Panha & J. B. Burch, 2004
Hypselostoma troglodytes van Benthem Jutting, 1950
Hypselostoma tubiferum Benson, 1856
Hypselostoma utongensis Panha & J. B. Burch, 2004^[6]
Hypselostoma venustum (van Benthem Jutting, 1950)
Hypselostoma vesovici Gojšina & Páll-Gergely, 2025
Hypselostoma vicinum Gojšina, Auffenberg & Páll-Gergely, 2025
Hypselostoma vujici Gojšina & Páll-Gergely, 2025

Synonyms

Hypselostoma dayanum Stoliczka, 1871: synonym of Acinolaemus dayanum (Stoliczka, 1871) (superseded combination)
Hypselostoma banmiensis Panha & J. B. Burch, 2004^[6]: synonym of Anauchen banmiensis Panha & J. B. Burch, 2004 (superseded combination)
Hypselostoma dohertyi Fulton, 1899: synonym of Hypselostoma everetti E. A. Smith, 1896 (junior subjective synonym)
Hypselostoma kentingensis C.-C. Hwang, 2014: synonym of Hypselostoma insularum Pilsbry, 1908 (junior subjective synonym)
Hypselostoma lacrima Páll-Gergely & Hunyadi, 2015^[7]: synonym of Clostophis lacrima (Páll-Gergely & Hunyadi, 2015) (> superseded combination)
Hypselostoma neglecta van Benthem-Jutting, 1961: synonym of Clostophis neglectus (van Benthem Jutting, 1961)
Hypselostoma rochebruni Mabille, 1887: synonym of Anauchen rochebruni (Mabille, 1887) (superseded combination)
Hypselostoma satulensis Panha & J. B. Burch, 2004^[6]: synonym of Hypselostoma hungerfordianum Möllendorff, 1891 (superseded combination)
Hypselostoma socialis Páll-Gergely & Hunyadi, 2015^[7]: synonym of Clostophis socialis (Páll-Gergely & Hunyadi, 2015) (superseded combination, original combination)
Hypselostoma striolatum (Möllendorff, 1895): synonym of Hypselostoma hungerfordianum Möllendorff, 1891 (junior subjective synonym)

References

^ Benson, W. H. (1856). "Remarks on the Genera Tanystoma, Nematura, and Anaulus". Ann. Mag. Nat. Hist. 2. 17 (100): 342. doi:10.1080/00222935608697520.
^ Solem, A. (1981). Small land snails from Northern Australia, I: Species of Gyliotrachela Tomlin, 1930 (Mollusca: Pulmonata, Vertiginidae). Journal of the Malacological Society of Australia 5(1-2) 87-100.
^ "WoRMS - World Register of Marine Species - Hypselostoma Benson, 1856". www.marinespecies.org. Retrieved 2020-01-21.
^ Vermeulen, Jaap; Luu, Hong Truong; Theary, Keum; Anker, Katja (2019-02-11). "New species of land snails (Mollusca: Gastropoda: Caenogastropoda and Pulmonata) of the Mekong Delta Limestone Hills (Cambodia, Vietnam)". Folia Malacologica. 27: 7–41. doi:10.12657/folmal.027.001.
^ Somsak, Panha; Burch, John B. (February 2002). "New Pupilloid Land Snails from Thailand (Pulmonata: Pupillidae)". The Natural History Journal of Chulalongkorn University. 2 (1): 21–24.
^ ^a ^b ^c ^d Panha, Somsak; Tongkerd, Piyoros; Sutjarit, Jirasak; Burch, John (2004-11-01). "New Pupillid Species from Thailand (Pulmonata: Pupillidae)". The Natural History Journal of Chulalongkorn University. 4: 57–82.
^ ^a ^b Páll-Gergely, Barna; Hunyadi, András; Jochum, Adrienne; Asami, Takahiro (2015-09-28). "Seven new hypselostomatid species from China, including some of the world's smallest land snails (Gastropoda, Pulmonata, Orthurethra)". ZooKeys (523): 31–62. doi:10.3897/zookeys.523.6114. ISSN 1313-2970. PMC 4602296. PMID 26478698.

External links

Gojšina, V., Hunyadi, A., Sutcharit, C., Tongkerd, P., Auffenberg, K., Grego, J., Vermeulen, J. J., Reischütz, A. & Páll-Gergely, B. (2025). "A new start? Revision of the genera Anauchen, Bensonella, Gyliotrachela and Hypselostoma (Gastropoda, Eupulmonata, Hypselostomatidae) of Southeast Asia with description of 46 new species". Zookeys. 1235. Retrieved 25 January 2026.{{cite journal}}: CS1 maint: multiple names: authors list (link)

[1] Benson, W. H. (1856). "Remarks on the Genera Tanystoma, Nematura, and Anaulus". Ann. Mag. Nat. Hist. 2. 17 (100): 342. doi:10.1080/00222935608697520.

[solem-2] Solem, A. (1981). Small land snails from Northern Australia, I: Species of Gyliotrachela Tomlin, 1930 (Mollusca: Pulmonata, Vertiginidae). Journal of the Malacological Society of Australia 5(1-2) 87-100.

[3] "WoRMS - World Register of Marine Species - Hypselostoma Benson, 1856". www.marinespecies.org. Retrieved 2020-01-21.

[4] Vermeulen, Jaap; Luu, Hong Truong; Theary, Keum; Anker, Katja (2019-02-11). "New species of land snails (Mollusca: Gastropoda: Caenogastropoda and Pulmonata) of the Mekong Delta Limestone Hills (Cambodia, Vietnam)". Folia Malacologica. 27: 7–41. doi:10.12657/folmal.027.001.

[5] Somsak, Panha; Burch, John B. (February 2002). "New Pupilloid Land Snails from Thailand (Pulmonata: Pupillidae)". The Natural History Journal of Chulalongkorn University. 2 (1): 21–24.

[:1-6] Panha, Somsak; Tongkerd, Piyoros; Sutjarit, Jirasak; Burch, John (2004-11-01). "New Pupillid Species from Thailand (Pulmonata: Pupillidae)". The Natural History Journal of Chulalongkorn University. 4: 57–82.

[:0-7] Páll-Gergely, Barna; Hunyadi, András; Jochum, Adrienne; Asami, Takahiro (2015-09-28). "Seven new hypselostomatid species from China, including some of the world's smallest land snails (Gastropoda, Pulmonata, Orthurethra)". ZooKeys (523): 31–62. doi:10.3897/zookeys.523.6114. ISSN 1313-2970. PMC 4602296. PMID 26478698.

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