2026 in reptile paleontology

Fossil reptile research published in 2026 includes peer-reviewed publications on discoveries related to reptile paleontology, as well as the description of new taxa.


Lepidosauromorphs

Squamates

Name Novelty Status Authors Age Type locality Country Notes Images

Paleoteius[1]

Gen. et sp. nov

Agnolín et al.

Late Cretaceous (Maastrichtian)

Allen Formation

Argentina

A possible member of Scincomorpha. The type species is P. lakui.

Pluridens imelaki[2]

Sp. nov

Longirch & Jalil

Late Cretaceous (Maastrichtian)

Upper Couche III

Morocco

A mosasaur belonging to the subfamily Halisaurinae; a species of Pluridens.

Prognathodon cipactli[3]

Sp. nov

Valid

Rivera-Sylva et al.

Late Cretaceous (Maastrichtian)

Méndez Shale

Mexico

Xenodontolacerta[4]

Gen. et sp. nov

Valid

Jiang et al.

Late Cretaceous (Coniacian–Campanian)

Hekou Formation

China

A polyglyphanodontian. The type species is X. fangi

Squamate research

  • Ebel, Melville & Keogh (2026) reconstruct the evolutionary history of squamate osteoderms on the basis of data from extant and extinct reptiles, reporting evidence of 13 independent acquisitions of osteoderms, the majority of which happened in the Late Jurassic and Early Cretaceous.[5]
  • Piñuela et al. (2026) report the discovery of lizard trackways from the Upper Jurassic (Kimmeridgian) strata of the Lastres Formation (Spain), interpreted as the latest occurrence of the ichnogenus Rhynchosauroides reported to date.[6]
  • Evidence indicating that inner ear morphology is an accurate predictor of higher-order classification of extant and fossil toxicoferans is presented by Forcellati et al. (2026).[7]
  • Xing et al. (2026) report evidence of a giant bone cell tumor within two digits of an anguimorph lizard specimen preserved within the Cretaceous Kachin amber (Myanmar).[8]
  • Redescription and a study on the affinities of Prognathodon waiparaensis is published by Young et al. (2026).[9]
  • Comans, Tobin & Totten (2026) reconstruct the thermoregulatory modes of Platecarpus and Tylosaurus from the Smoky Hill Chalk Member of the Niobrara Formation (Kansas, United States) on the basis stable oxygen isotope composition of tooth enamel, interpreted as consistent with endothermy.[10]
  • Datta & Bajpai (2026) report the discovery of new fossil material of constrictor snakes from the Ypresian strata of the Cambay Shale and from the Lutetian strata from Panandhro (India), possibly representing new taxa, and preserving evidence of differences of body size of snakes from the two sites which might be indicative of ecological and environmental differences.[11]
  • Liaw et al. (2026) report the discovery of a vertebra of a large-bodied python from the Pleistocene strata of the Chiting Formation (Taiwan).[12]
  • Jansen et al. (2026) report the discovery of a new assemblage of squamate fossils from the Campanian strata of the Villeveyrac-Mèze basin (France), including the oldest European members of Pan-Shinisaurus, Madtsoiidae, Monstersauria and Iguanomorpha reported to date, and possibly the oldest known anguid worldwide.[13]

Other lepidosauromorphs

Name Novelty Status Authors Age Type locality Country Notes Images

Airistagiz[14]

Gen. et sp. nov

Valid

Sobral & Schoch

Middle Triassic (Ladinian)

Erfurt Formation

Germany

A stem-lepidosaur. The type species is A. seegisi.

Hohlachia[14]

Gen. et sp. nov

Valid

Sobral & Schoch

Middle Triassic (Ladinian)

Erfurt Formation

Germany

A stem-lepidosaur. The type species is H. multidens.

Klainjosaura[14]

Gen. et sp. nov

Valid

Sobral & Schoch

Middle Triassic (Ladinian)

Erfurt Formation

Germany

A stem-lepidosaur. The type species is K. staroskalja.

Other lepidosauromorph research

  • Haridy et al. (2026) describe new fossil material of Eilenodon robustus from the strata of the Morrison Formation from Utah and Wyoming (United States), and provide the first three-dimensional reconstructions of the skull anatomy of this taxon and the first detailed study of its tooth row histology.[15]
  • Cavasin, Cerda & Apesteguía (2026) study the histology of the beak-like structure in Priosphenodon avelasi, and report that the studied structure is not formed by teeth fused to the premaxillae, but instead it is entirely formed by bone tissue.[16]

Archosauromorphs

Archosaurs

Main article: 2026 in archosaur paleontology

Other archosauromorphs

Name Novelty Status Authors Age Type locality Country Notes Images

Other archosauromorph research

  • Sarkar & Ray (2026) report evidence from the study of the bone histology of members of an assemblage of Colossosuchus techniensis from the Tiki Formation (India) indicative of an epidemic of persistent, recurrent bone disease in members of the studied community, likely resulting from a bacterial infection.[17]

Turtles

Name Novelty Status Authors Age Type locality Country Notes Images

Gigantochelys[18]

Gen. et sp. nov

Mousa, Tantawy & El-Kheir

Late Cretaceous (Maastrichtian)

Dakhla Formation

Egypt

A member of the family Dermochelyidae. The type species is G. aegyptiacus.

Sternotherus pugnatus[19]

Sp. nov

Valid

Bourque

Miocene

United States
( Florida)

A species of Sternotherus.

Turtle research

  • Pérez-García et al. (2026) revise Neochelys arribasi, and support its recognition as a valid species.[20]
  • Pochat-Cottilloux et al. (2026) report the first discovery of trionychid fossil material (including Trionyx cf. vindobonensis) from the Miocene strata of Poland, interpreted as indicative of environmental conditions suitable for animals requiring high temperatures between the Middle Miocene Climatic Optimum and Late Miocene in the studied area.[21]
  • Footprint traces interpreted as likely produced by a stampede of sea turtles panicked by an earthquake are reported from the Campanian strata from Monte Cònero (Italy) by Sandroni et al. (2026).[22]

Other reptiles

This section includes reptiles of uncertain or debated placement (choristoderes and the marine reptile clades Ichthyosauromorpha and Sauropterygia) in addition to stem group reptiles.

Choristoderes

Choristodere research

  • Matsumoto, Manabe & Evans (2026) describe new fossil material of members of Choristodera from the Lower Cretaceous Okurodani Formation (Japan), expanding known diversity of members of the group from the studied formation.[23]

Ichthyosauromorphs

Name Novelty Status Authors Age Type locality Country Notes Images

Ichthyosauromorph research

  • A study on the anatomy of the ichthyopterygian specimen NSM-PV-20028 from the Lower Triassic Osawa Formation (Japan), assigned to cf. Utatsusaurus hataii, is published by Yoshizawa & Tsuihiji (2026).[24]
  • Evidence indicating that the relative size of the notochordal canal on vertebral centra can be used to study developmental stage of ichthyosaur fetuses is presented by Miedema & Maxwell (2026).[25]
  • Probable ophthalmosaurid specimen representing the first partially articulated ichthyosaur skeleton reported from the insular Caribbean is described from the Tithonian strata of the Guasasa Formation (Cuba) by Iturralde-Vinent et al. (2026).[26]
  • Martinez-Motta et al. (2026) report evidence of preservation of integumentary tissues in a brachypterygiid ichthyosaur specimen from the Paja Formation (Colombia).[27]

Sauropterygians

Name Novelty Status Authors Age Type locality Country Notes Images

Gondwananectes[28]

Gen. et sp. nov

Otero et al.

Middle Jurassic (Bajocian)

Quehuita Formation

Chile

A plesiosaur with affinities with Cryptoclidia. The type species is G. osvaldoi.

Nothosaurus fortihumeralis[29] Sp. nov. Li et al. Middle Triassic (Anisian) Guanling Formation China A nothosaurid

Pahasapasaurus gillettei[30]

Sp. nov.

In press

Schmeisser McKean

Late Cretaceous (Turonian)

Tropic Shale

United States ( Utah)

A polycotylid plesiosaur; a species of Pahasapasaurus. Announced in 2025, the final article version will be published in 2026.

Sauropterygian research

  • Zhang et al. (2026) determine the age and duration of the Ladinian Xingyi Fauna on the basis of the study of astrochronology and cyclostratigraphy of the Nimaigu Section of the Falang Formation (China), providing evidence of brief duration of the typical species of this fauna, Keichousaurus hui.[31]
  • Redescription of the anatomy and a study on the affinities of Lusonectes sauvagei is published by Sachs & Madzia (2026).[32]
  • A study on the anatomy and affinities of "Plesiosaurus" posidoniae is published by Sachs, Schweigert & Madzia (2026).[33]
  • García-Guerrero et al. (2026) describe a cervical vertebra of a member of the subfamily Brachaucheninae from the Valanginian strata of the Rosablanca Formation (Colombia), representing the oldest fossil material of a large pliosaurid from the Lower Cretaceous strata in northern South America reported to date.[34]
  • Probable elasmosaurid vertebra, representing the first plesiosaur record from Algeria, is described from the Coniacian strata of the Essen Formation in the Tébessa Mountains by Nadir Naimi et al. (2026).[35]
  • Evidence of a healing fracture and periostitis is reported in elasmosaurid specimens from the Maastrichtian Snow Hill Island Formation (Antarctica) and Jagüel Formation (Argentina) by Mitidieri et al. (2026).[36]
  • Marx, Szasz & Lindgren (2026) determine heat transfer in elasmosaurid models reconstructed with and without a layer of insulating blubber, and argue that a peripheral blubber layer was necessary for elasmosaurids inhabiting cold water regions.[37]
  • Drumheller et al. (2026) report the discovery of a fish tooth embedded in a cervical vertebra of a specimen of Polycotylus latipinnis from the Cretaceous Mooreville Chalk (Alabama, United States), interpreted as likely evidence of an attack by Xiphactinus.[38]

Stem group reptiles

Name Novelty Status Authors Age Type locality Country Notes Images

Sauropia[39]

Gen. et sp. nov

Valid

Müller et al.

Middle Triassic (Ladinian)

Pinheiros-Chiniquá Sequence of the Santa Maria Supersequence

Brazil

A member or relative of the family Procolophonidae. The type species is S. macrorhinus.

Scyllacerta[40] Gen. et sp. nov Valid Jenkins et al. Late Permian Teekloof Formation (Endothiodon Assemblage Zone) South Africa A younginid. The type species is S. creanae.

Stem group reptile research

  • Marchetti et al. (2026) report the discovery of resting trace with a probable bolosaurian skin impression from the Permian (Asselian) Goldlauter Formation (Germany), including a probable cloacal vent impression and the oldest definite impression of epidermal scales of a reptile reported to date, and name a new ichnotaxon Cabarzichnus pulchrus.[41]
  • D'Ávila Burgardt et al. (2026) report the discovery of a new procolophonid lower jaw from the Lower Triassic Sanga do Cabral Formation (Brazil), differing from known procolophonid taxa from the site in dental morphology.[42]

Reptiles in general

  • Evidence from the study of body size of reptiles from the Shungura Formation (Ethiopia), indicative of temporal coincidence between changes of maximum body size of members of the studied assemblage and local environmental changes (including changes in hydrological regimes and evaporation levels), is presented by Parker et al. (2026).[43]
  • A second specimen of Yaverlandia bitholus is described by Naish & Sweetman (2026), who consider the dinosaurian affinities of the taxon to be questionable.[44]

References

  1. ^ Agnolín, F. L.; Aranciaga-Rolando, M.; Álvarez-Herrera, G.; Ezcurra, M. D.; Moreno Rodríguez, A.; Chafrat, P.; Vega, N.; Scanferla, A.; Smith, K. T.; Novas, F. E. (2026). "A new late Cretaceous squamate from Patagonia sheds light on Gondwanan diversity". Scientific Reports. doi:10.1038/s41598-026-40914-8. PMID 41764267.
  2. ^ Longrich, Nicholas R.; Nour-Eddine, Jalil (2026-03-05). "A Giant Halisaurine from the Late Maastrichtian of Morocco". Diversity. 18 (3) 159. doi:10.3390/d18030159.
  3. ^ Rivera-Sylva, H. E.; Sánchez-Uribe, I. E.; Guzmán-Gutierrez, J. R.; Rodríguez, R. L. N.; Rangel-Morelos, A. A.; Longrich, N. (2026). "A new species of Prognathodon (Squamata: Mosasauridae) from the Early Maastrichtian of Nuevo León, Mexico". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. doi:10.1127/njgpa/1303.
  4. ^ Jiang, Juan; Dong, Li-Ping; Xu, Xing; Bi, Alexander; Evans, Susan (2026-02-04). "A new polyglyphanodontian lizard from the Upper Cretaceous of southern China implies a complex evolutionary history of the clade". Royal Society Open Science. 13 (2). doi:10.1098/rsos.252253. ISSN 2054-5703.
  5. ^ Ebel, R.; Melville, J.; Keogh, J. S. (2026). "Lizards in chain mail: reconstructing the enigmatic past of dermal armour in squamate reptiles". Biological Journal of the Linnean Society. 147 (1) blaf129. doi:10.1093/biolinnean/blaf129.
  6. ^ Piñuela, L.; Xing, L.; García-Pérez, Á.; García-Ramos, J. C. (2026). "First report of lizard trackways from the Jurassic of Europe". Ichnos. doi:10.1080/10420940.2026.2634103.
  7. ^ Forcellati, M. R.; Napoli, J. G.; Meyer, D.; Watanabe, A.; Benson, R.; Raxworthy, C. J. (2026). "Machine learning can accurately assign fossil and extant species to crown toxicoferan (Reptilia: Squamata) groups using inner ear shape data". Zoological Journal of the Linnean Society. 206 (3) zlaf188. doi:10.1093/zoolinnean/zlaf188.
  8. ^ Xing, L.-D.; Rothschild, B. M.; Li, J.-L.; Peng, A.-C.; Jin, Y.-F.; Liu, X.-R.; McKellar, R. (2026). "A bone lesion in a lizard from mid-Cretaceous Burmese amber". Palaeoentomology. 9 (1): 85–98. doi:10.11646/palaeoentomology.9.1.10.
  9. ^ Young, G.; Mannering, A.; Fraser, M.; Scofield, P. (2026). "Redescription of Prognathodon waiparaensis and comments on its phylogenetic position". Alcheringa: An Australasian Journal of Palaeontology. doi:10.1080/03115518.2025.2607629.
  10. ^ Comans, C. M.; Tobin, T. S.; Totten, R. L. (2026). "Evidence for endothermy from tooth enamel(oid) oxygen isotopes in marine predators of the Late Cretaceous Western Interior Seaway, USA". Palaeogeography, Palaeoclimatology, Palaeoecology 113578. doi:10.1016/j.palaeo.2026.113578.
  11. ^ Datta, D.; Bajpai, S. (2026). "Eocene constrictors (Serpentes; Alethinophidia) from western India: New taxonomic and paleobiogeographic insights". The Anatomical Record. doi:10.1002/ar.70174.
  12. ^ Liaw, Y.-L.; Cho, Y.-Y.; Sun, C.-H.; Biswas, D. S.; Tsai, C.-H. (2026). "An unexpected snake fossil (Pythonidae, Python) from Taiwan". Historical Biology: An International Journal of Paleobiology. doi:10.1080/08912963.2025.2610741.
  13. ^ Jansen, O.; Garcia, G.; Otero, O.; Augé, M.; Gomez, B.; Valentin, X. (2026). "Freshwater amphibians and squamates from Villeveyrac (lower Campanian; Hérault, France): palaeodiversity, palaeoenvironment and implications for the Late Cretaceous palaeobiogeography of the European herpetofauna". Papers in Palaeontogy. 12 (1) e70055. doi:10.1002/spp2.70055.
  14. ^ a b c Sobral, Gabriela; Schoch, Rainer R. (2026-02-19). "New stem lepidosaurs from Vellberg, Germany: implications for palaeoecology in the early diversification of Lepidosauromorpha". Journal of Systematic Palaeontology. 24 2603472. doi:10.1080/14772019.2025.2603472.
  15. ^ Haridy, Y.; Frederickson, J. A.; Peterson, J.; Miller, J.; Davis, B.; Curtice, B.; Hunter, R.; Cifelli, R. (2026). "New material of the Late Jurassic sphenodontid Eilenodon robustus reveals transitionary adaptations to herbivory". New Mexico Museum of Natural History and Science Bulletin. 102: 171–184.
  16. ^ Cavasin, S. A.; Cerda, I. A.; Apesteguía, S. (2026). "Bone microstructure of the beak-like structure in the herbivorous sphenodontian Priosphenodon avelasi (Lepidosauria)". Cretaceous Research 106363. doi:10.1016/j.cretres.2026.106363.
  17. ^ Sarkar, P.; Ray, S. (2026). "Extensive radial fibrolamellar bone tissue in long bones reveals an epidemic of persistent, recurrent, and non-traumatic bone infection in a juvenile-dominated phytosaur (Archosauria; Pseudosuchia) community". The Anatomical Record. doi:10.1002/ar.70160. PMID 41708148.
  18. ^ Mousa, M. K.; Tantawy, A. A. A.; El-Kheir, G. A. A. (2026). "First known gigantic dermochelyid turtle (Pan-Chelonioidea) from the Upper Cretaceous deposits of Dakhla Oasis, Western Desert, Egypt". Cretaceous Research 106357. doi:10.1016/j.cretres.2026.106357.
  19. ^ Bourque, J. R. (2026). "A small flat-shelled musk turtle from the Late Miocene of Florida and new Pleistocene records of Sternotherus (Kinosternidae)". Historical Biology: An International Journal of Paleobiology. doi:10.1080/08912963.2025.2602686.
  20. ^ Pérez-García, A.; Guerrero, A.; Martín de Jesús, S.; Ortega, F. (2026). "Evaluation of the Validity of the Spanish Eocene Pleurodiran Turtle 'Duerochelys arribasi': The Youngest Freshwater Member of Erymnochelyinae (Podocnemididae) in Europe". Diversity. 18 (1) 51. doi:10.3390/d18010051.
  21. ^ Pochat-Cottilloux, Y.; López-Torres, S.; Chroust, M.; Georgalis, G. L.; Górka, M.; Tałanda, M. (2026). "First occurrences of Trionychidae (Testudines, Cryptodira) from the Miocene of Poland: Detailed cranial anatomy and biogeographic implications". Journal of Anatomy. doi:10.1111/joa.70095.
  22. ^ Sandroni, P.; Church, N. S.; Coccioni, R.; Frontalini, F.; Mainiero, M.; Montanari, A. (February 2026). "Reptile footprints on a pelagic seafloor as a vestige of a synsedimentary seismic event in the lower Campanian Scaglia Rossa Basin of the Umbria-Marche Apennines (Italy)". Cretaceous Research. 179 106268. doi:10.1016/j.cretres.2025.106268.
  23. ^ Matsumoto, R.; Manabe, M.; Evans, S. E. (May 2026). "Cranial elements of Shokawa ikoi, and new records of choristoderes from the Lower Cretaceous Okurodani Formation, Tetori Group, Japan". Cretaceous Research. 181 106283. doi:10.1016/j.cretres.2025.106283.
  24. ^ Yoshizawa, K.; Tsuihiji, T. (2026). "Skeletal morphology of an early ichthyopterygian, cf. Utatsusaurus hataii, from Japan with an emphasis on CT scan data of the skull". Journal of Vertebrate Paleontology e2621687. doi:10.1080/02724634.2026.2621687.
  25. ^ Miedema, F.; Maxwell, E. E. (2026). "Use of the notochordal canal as a reliable proxy for prenatal stage, a case study in Ichthyosauria". Royal Society Open Science. 13 (2) 251986. doi:10.1098/rsos.251986.
  26. ^ Iturralde-Vinent, M.; Campos, L.; Pszczółkowski, A.; Ceballos-Izquierdo, Y.; Viñola-Lopez, L. W. (2026). "A partial ichthyosaur (?Ophthalmosauridae) skeleton from the Tithonian (Upper Jurassic) of western Cuba". Journal of Vertebrate Paleontology e2609717. doi:10.1080/02724634.2025.2609717.
  27. ^ Martinez-Motta, M. F.; Cortés, D.; Maxwell, E. E.; Noè, L. F.; Alfonso-Rojas, A.; Parra-Ruge, F. H.; Cadena, E.-A. (2026). "Soft-tissue preservation in an ichthyosaur from the Lower Cretaceous (Barremian - Aptian) of Colombia". Cretaceous Research 106305. doi:10.1016/j.cretres.2025.106305.
  28. ^ Otero, R. A.; Soto Acuña, S.; Vargas, A. O.; Rojas, J.; Ortiz, H.; Aguirrezabala, G. (2026). "A new Middle Jurassic marine reptile from Gondwana clarifies the origin of Cryptoclidia, the most successful group of plesiosaurs". Papers in Palaeontology. 12 (2) e70068. doi:10.1002/spp2.70068.
  29. ^ Li, Qiang; Hu, Yi-Wei; Shen, Yuefeng; Sander, P Martin; Liu, Jun (2026-01-01). "A new large Nothosaurus (Reptilia: Eosauropterygia) from the Middle Triassic of South China and the phylogeny of Eosauropterygia". Zoological Journal of the Linnean Society. 206 (1). doi:10.1093/zoolinnean/zlaf179. ISSN 0024-4082.
  30. ^ Schmeisser McKean, Rebecca L. (2026). "A new species of Pahasapasaurus (Plesiosauria: Polycotylidae) from the Upper Cretaceous Tropic Shale (lower Turonian) of southern Utah, U.S.A.". Cretaceous Research. 180 106269. doi:10.1016/j.cretres.2025.106269.
  31. ^ Zhang, Y.; Jiang, D.; Motani, R.; Tintori, A.; Sun, Z.; Zhou, M.; Su, C. X.; Wang, Y.; Yao, M. (2026). "Astronomical age calibration and duration of the Ladinian (Middle Triassic) Xingyi Fauna, southwestern Guizhou Province, China". Palaeogeography, Palaeoclimatology, Palaeoecology. 689 113652. doi:10.1016/j.palaeo.2026.113652.
  32. ^ Sachs, S.; Madzia, D. (2026). "The osteology, taxonomy, and phylogenetic affinities of the Early Jurassic plesiosaur Lusonectes sauvagei". PeerJ. 14 e20611. doi:10.7717/peerj.20611. PMC 12875220.
  33. ^ Sachs, S.; Schweigert, G.; Madzia, D. (2026). "Unraveling the identity of Plesiosaurus posidoniae, the first plesiosaur described from the Posidonia Shale of Germany". The Anatomical Record. doi:10.1002/ar.70157.
  34. ^ García-Guerrero, J.; Parra Ruge, F.; Prieto, G. A.; Cadena, E.-A. (February 2026). "Evidence of large pliosaurids in the Late Valanginian of Colombia". Cretaceous Research. 178 106235. doi:10.1016/j.cretres.2025.106235.
  35. ^ Nadir Naimi, M.; Salmi-Laouar, S.; O'Gorman, J. P.; Nemouchi, S.; Degaichia, A. (2026). "First plesiosaurian (Diapsida; Sauropterygia) remains from Algeria". Historical Biology: An International Journal of Paleobiology. doi:10.1080/08912963.2026.2612987.
  36. ^ Mitidieri, M.; Talevi, M.; Herrera, Y.; Rothschild, B.; Fernández, M. S. (January 2026). "From the inside: pathologies from a histological perspective of two elasmosaurids from the Upper Cretaceous of Antarctica and Argentina". Cretaceous Research. 177 106208. doi:10.1016/j.cretres.2025.106208.
  37. ^ Marx, M.; Szasz, R.-Z.; Lindgren, J. (2026). "CFD modelling of the thermo- and hydrodynamic capabilities of long-necked plesiosaurs (Sauropterygia, Elasmosauridae)". Technische Mechanik - European Journal of Engineering Mechanics. 46 (1): 43–54. doi:10.24352/UB.OVGU-2026-006.
  38. ^ Drumheller, S. K.; O'keefe, F. R.; Mayhall, M. L.; Stalker, E.; Brochu, C. A. (2026). "A bite to the throat: A probable Xiphactinus attack on a Polycotylus from the Cretaceous Mooreville Chalk of Alabama, U.S.A." Journal of Vertebrate Paleontology e2625732. doi:10.1080/02724634.2026.2625732.
  39. ^ Müller, R. T.; Roberto-da-Silva, L.; Aurélio, P. L. P.; Kerber, L. (2026). "The smallest tetrapod from the Middle Triassic of South America: a new procolophonoid parareptile from the Ladinian of Southern Brazil". Scientific Reports. 16 866. doi:10.1038/s41598-026-35114-3. PMC 12852135. PMID 41606015.
  40. ^ Jenkins, Xavier A.; Buffa, Valentin; Marchant, Cy J.; Ford, David P.; Browning, Claire; Fernandez, Vincent; Dollman, Kathleen; Botha, Jennifer; Choiniere, Jonah N.; Benson, Roger B. J.; Peecook, Brandon R. (2026-01-22). "The origin of the tympanic fossa in reptiles revealed by a late Permian neodiapsid". Palaeontology. 69 (1) e70041. doi:10.1111/pala.70041. ISSN 0031-0239.
  41. ^ Marchetti, L.; Logghe, A.; Buchwitz, M.; MacDougall, M. J.; Rebillard, A.; Martens, T.; Fröbisch, J. (2026). "The earliest reptile body impressions with scaly skin". Current Biology. doi:10.1016/j.cub.2026.01.036. PMID 41687603.
  42. ^ D'Ávila Burgardt, C.; Pohlmann, K.; Machado, A. F.; Costa Santos, M. A.; Meira, J. L.; Pinheiro, F. L. (2026). "A peculiar procolophonid lower jaw from the Sanga do Cabral Formation (Lower Triassic, Brazil)". Historical Biology: An International Journal of Paleobiology. doi:10.1080/08912963.2025.2608698.
  43. ^ Parker, A. K.; Boisserie, J.-R.; Müller, J.; Brochu, C. A.; Head, J. J. (2026). "Body size histories of Shungura Formation reptiles in biotic and abiotic environmental context". Comptes Rendus Palevol. 25 (2): 17–42. doi:10.5852/cr-palevol2026v25a2.
  44. ^ Naish, Darren; Sweetman, Steven C. (2026-03-16). "A second specimen of the enigmatic Wealden reptile Yaverlandia". Proceedings of the Geologists' Association 101179. doi:10.1016/j.pgeola.2026.101179.
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