Stegophilinae

Stegophilinae
Ochmacanthus reinhardtii
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Siluriformes
Family: Trichomycteridae
Subfamily: Stegophilinae
Günther, 1864[1]
Type genus
Stegophilus[1]

Stegophilinae is a subfamily of freshwater ray-finned fishes belonging to the family Trichomycteridae, the pencil and parasitic catfishes. Stegophilines are sometimes considered candirú due to their parasitic or semiparasitic habits, because of their peculiar habit of feeding on scales, mucus, or the skin of other fishes.[3][4]

The Stegophilinae are widely distributed in the main South American river basins, including the Amazon, Orinoco, São Francisco, Paraná-Paraguay, and those of southern Brazil.[5] Eight of the genera are distributed in Venezuela.[3]

Taxonomy

The subfamily is defined by a few characters, such as osteological synapomorphies of the mandibular and suspensorial arches,[4] and a wide, crescent-shaped, non-occluding mouth, though Pareiodon "secondarily reversed" this trait and has differing mouth anatomy.[6]

Stegophilinae contains the following valid genera:[2]

Stegophilinae has consistently been found to form a monophyletic group (clade); the subfamily also encompass smaller clades which are supported by two characteristics of the lateral line, including Acanthopoma, Henonemus, Megalocentor, Pareiodon, Parastegophilus, and Pseudostegophilus. Acanthopoma and Henonemus were found to have a sister group relationship.[3] The following cladogram is based on a 2026 phylogenetic study of morphology using maximum parsimony:[4]

Tridentinae

References

  1. ^ a b Richard van der Laan; William N. Eschmeyer & Ronald Fricke (2014). "Family-group names of recent fishes". Zootaxa. 3882 (2): 1–230. doi:10.11646/zootaxa.3882.1.1. PMID 25543675.
  2. ^ a b Fricke, Ron; Eschmeyer, William N. & van der Laan, Richard (eds.). "Genera in the family Stegophilinae". Catalog of Fishes. California Academy of Sciences. Retrieved 19 November 2025.
  3. ^ a b c DoNascimiento, Carlos; Provenzano, Francisco (2006). "The Genus Henonemus (Siluriformes: Trichomycteridae) with a Description of a New Species from Venezuela". Copeia. 2006 (2): 198–205. doi:10.1643/0045-8511(2006)6[198:TGHSTW]2.0.CO;2. S2CID 84355922.
  4. ^ a b c "Phylogenetic assessment and taxonomic description of the first scale-feeding candiru from west of the Andes (Siluriformes, Trichomycteridae)". Zoological Journal of the Linnean Society. 206 (3). 15 March 2026. doi:10.1093/zoolinnean/zlag002.
  5. ^ Koch, Walter Rudolf (30 September 2002). "Revisão Taxonômica do Gênero Homodiaetus (Teleostei, Siluriformes, Trichomycteridae) (Portuguese)" (PDF). Iheringia, Sér. Zool., Porto Alegre. 92 (3): 33–46. doi:10.1590/s0073-47212002000300004.
  6. ^ DoNascimiento, C.; Monophyly of the subfamily of parasitic catfishes Stegophilinae (Siluriformes, Trichomycteridae) and phylogenetic diagnoses of its genera. Copeia 2015; 103: 933–60. DOI