The class Notopharyngea, formerly known as the BaSk clade, includes two orders of free-living anaerobic metamonads. It was identified by phylogenomic analyses as a deeply branching sister group to the Fornicata, and recently the rest of Metamonada. The extreme mitochondrial reduction observed within this free-living anaerobic protistan clade demonstrates that mitochondrial functions may be completely lost even in free-living organisms, not only parasites.[1][2]
References
- ^ Williams, Shelby K.; Jerlström Hultqvist, Jon; Eglit, Yana; Salas-Leiva, Dayana E.; Curtis, Bruce; Orr, Russell J. S.; Stairs, Courtney W.; Atalay, Tuğba N.; MacMillan, Naomi; Simpson, Alastair G. B.; Roger, Andrew J. (2024-08-09). "Extreme mitochondrial reduction in a novel group of free-living metamonads". Nature Communications. 15 (1): 6805. doi:10.1038/s41467-024-50991-w. ISSN 2041-1723. PMC 11316075. PMID 39122691.
- ^ Shiratori, Takashi; Eglit, Yana; Yazaki, Euki; Inagaki, Yuji; Simpson, Alastair G. B. (2026-04-03). "Morphology, Ultrastructure, and Classification of Barthelonids (Metamonada Incertae Sedis): With Descriptions of Microbarthelona Gen. Nov., and Parabarthelona Gen. Nov". Journal of Eukaryotic Microbiology. 73 (3). doi:10.1111/jeu.70080. ISSN 1066-5234. PMC 13087550. PMID 41992732.
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- Domain
- Archaea
- Bacteria
- Eukaryota
- (major groups
- Metamonada
- Discoba
- Diaphoretickes
- Hacrobia
- Cryptista
- Rhizaria
- Alveolata
- Stramenopiles
- Plants
- Amorphea
- Amoebozoa
- Opisthokonta
- Animalia
- Fungi
- Mesomycetozoea)
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- Chilomitus?
- Rhizonympha?
- Tricercomitus?
- Trichocovina?
| Hypotrichomonadea | | Hypotrichomonadida |
- Hypotrichomonadidae
- Hypotrichomonas
- Trichomitus
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| Pimpavickea | |
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| Trichomonadea | | Honigbergiellida |
- Hexamastigidae
- Honigbergiellidae
- Honigbergiella
- Ditrichomonoides
- Honigbergiellopsis
- Monotrichomonas
- Cthulhuidae
- Tetratrichomastigidae
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| Trichomonadida | |
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Clade "Tla" | | Lophomonadea | | Lophomonadida |
- Lophomonadidae
- Lophomonas
- Eulophomonas
- Prolophomonas
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| Trichonymphea | | Trichonymphida |
- Barbulanymphidae
- Hoplonymphidae
- Retractinymphidae
- Spirotrichosomidae
- Spirotrichosoma
- Apospironympha
- Bispironympha
- Colospironympha
- Heliconympha
- Leptospironympha
- Macrospironympha
- Staurojoeninidae
- Teranymphidae
- Teranympha
- Eucomonympha
- Pseudotrichonympha
- Trichonymphidae
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| Clade "Cadamassta" | | Cristamonadea |
- Cyclojoenia?
- Joenia?
- Joenina?
- Joenoides?
- Joenopsis?
- Pachyjoenia?
- Parajoenopsis?
- Placojoenia?
- Projoenia?
| Calonymphida |
- Calonymphidae
- Calonympha
- Criconympha
- Daimonympha
- Diplonympha
- Gyronympha
- Metastephanonympha
- Prosnyderella
- Snyderella
- Stephanonympha
- Deltotrichonymphidae
- Mixotrichidae
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| Devescovinida |
- Devescovinidae
- Devescovina
- Achemon
- Astronympha
- Bullanympha
- Caduceia
- Coronympha
- Evemonia
- Foaina
- Hyperdevescovina
- Kirbynia
- Kofoidia
- Macrotrichomonas
- Macrotrichomonoides
- Metadevescovina
- Parajoenia
- Polymastigoides
- Pseudodevescovina
- Runanympha
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| Gigantomonadida | |
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| Dientamoebea | |
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| Monocercomonadea | |
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| Simplicimonadea | |
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| Tritrichomonadea | |
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| Spirotrichonymphea | | Cononymphida | |
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| Holomastigotoidida | |
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| Spirotrichonymphida |
- Brugerollinidae
- Brugerollina
- Pseudospironympha
- Fraterculidae
- Holomastigotidae
- Holomastigotes
- Spiromastigotes
- Spirotrichonymphella
- Uteronympha
- Spirotrichonymphidae
- Spirotrichonympha
- Microjoenia
- Micromastigotes
- Nanospironympha
- Spironympha
- Torquenympha
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