List of Ramalina species

This is a list of species in Ramalina, a genus of fruticose lichens. As of February 2026, Species Fungorum (in the Catalogue of Life) accepts 281 species of Ramalina.[1]

A

  • Ramalina abyssinica Nyl. (1870)[2]
  • Ramalina ahtii Kashiw. & T.H.Nash (1994)[3] – Mexico
  • Ramalina ailaoshanensis S.Y.Guo & L.F.Han (2021)[4] – China
  • Ramalina alisiosae Pérez-Vargas & Pérez-Ort. (2013)[5] – Canary Islands
  • Ramalina americana Hale (1979)[6] – North America
  • Ramalina andina V.Marcano & A.Morales (1994)[7] – Venezuela
  • Ramalina anisitsiana Gyeln. (1931)
  • Ramalina anteojina A.Morales & V.Marcano (2021)
  • Ramalina arabum (Dill. ex Ach.) Meyen & Flot. (1843)
  • Ramalina arbuscula Stizenb. (1890)
  • Ramalina arsenii Sérus., van den Boom & Magain (2021)[8] – Europe
  • Ramalina asahinana Zahlbr. (1927)
  • Ramalina aspera Räsänen (1944)
  • Ramalina asperula Kremp. (1876)
  • Ramalina aulota Stirt. (1874)
  • Ramalina australiensis Nyl. (1870)[2]
  • Ramalina azorica Aptroot & Schumm (2008)[9] – Azores

B

  • Ramalina banzarensis C.W.Dodge (1948)[10]
  • Ramalina bermudana Stirt. (1874)[11]
  • Ramalina bicolor Müll.Arg. (1877)[12]
  • Ramalina bigeniculata B.de Lesd. (1913)[13]
  • Ramalina bistorta Nyl. (1870)[2]
  • Ramalina boninensis Asahina (1938)[14]
  • Ramalina boulhautiana Maheu & A.Gillet (1925)
  • Ramalina bourgaeana Mont. ex Nyl. (1870)[2]
  • Ramalina brevis F.Wilson (1889)
  • Ramalina breviuscula (Nyl.) Nyl. (1872)[15]

C

  • Ramalina caespitella G.N.Stevens (1986)[16] – Australia
  • Ramalina caespitosa H.Magn. (1955)
  • Ramalina calcarata Krog & Swinscow (1974)[17] – East Africa
  • Ramalina calicaris (L.) Röhl. (1813)
  • Ramalina camptospora Nyl. (1870)[2]
  • Ramalina canaguensis V.Marcano & A.Morales (1994)[18]
  • Ramalina canalicularis (Nyl.) Kashiw. (2004)
  • Ramalina canariensis J.Steiner (1904)[19]
  • Ramalina cannonii Elix, Laily & Samsudin (1991)[20] – Peninsular Malaysia
  • Ramalina capensis Th.Fr. (1861)
  • Ramalina capitata (Ach.) Nyl. (1872)[15]
  • Ramalina caracasana Müll.Arg. (1877)[12]
  • Ramalina carminae R.Arroyo & Seriñá (2011)
  • Ramalina celastri (Spreng.) A.Massal. (1861)[21]
  • Ramalina ceratea Werner (1954)
  • Ramalina cerinella Zahlbr. (1945)
  • Ramalina cerinelloides H.Magn. (1955)
  • Ramalina chiguarensis V.Marcano & A.Morales (1994)[7] – Venezuela
  • Ramalina chihuahuana Kashiw. & T.H.Nash (2002)[22] – Mexico
  • Ramalina chondrina J.Steiner (1904)[19]
  • Ramalina cinereovirens Kashiw., K.H.Moon & J.E.Han (2021)[23] – South Korea
  • Ramalina clementeana Llimona & Werner (1975)[24]
  • Ramalina cochlearis Zahlbr. (1905)[25]
  • Ramalina commixta Asahina (1939)
  • Ramalina conduplicans Vain. (1921)[26]
  • Ramalina confirmata (Nyl.) Zahlbr. (1930)
  • Ramalina confusa D.D.Awasthi (2006)
  • Ramalina congesta Kashiw., K.H.Moon & J.E.Han (2023)[27] – Japan, Korea
  • Ramalina continentalis Malme (1935)
  • Ramalina controversa Sbarbaro (1955)
  • Ramalina coreana Kashiw. & K.H.Moon (2002)[28] – Southeast Asia
  • Ramalina corsicana Zahlbr. (1927)
  • Ramalina corymbosa (Hue) Kotlov (2004)[29]
  • Ramalina costata Meyen & Flot. (1843)[30]
  • Ramalina cribrosa De Not. (1846)[31]
  • Ramalina crispata V.Marcano & A.Morales (1994)[7] – Venezuela
  • Ramalina culbersoniorum LaGreca (2000)[32]
  • Ramalina cumanensis Fée (1825)[33]
  • Ramalina cuspidata (Ach.) Nyl. (1870)[2]

D

  • Ramalina dalmatica J.Steiner & Zahlbr. (1903)
  • Ramalina darwiniana Aptroot & Bungartz (2007)[34] – Galapagos
  • Ramalina dasypoga Tuck. (1859)
  • Ramalina debrecenensis Gyeln. (1932)
  • Ramalina deliblatensis Gyeln. (1932)
  • Ramalina delicata Blázquez, Pérez-Vargas & Pérez-Ort. (2024)
  • Ramalina dendriscoidella Vain. (1926)
  • Ramalina dendriscoides Nyl. (1876)
  • Ramalina desolata H.Magn. (1942)
  • Ramalina dichotoma Hepp (1854)
  • Ramalina dictyota C.W.Dodge & Vareschi (1956)
  • Ramalina digitata Meyen & Flot. (1843)
  • Ramalina dilacerata (Hoffm.) Hoffm. (1825)
  • Ramalina disciformis Kashiw., K.H.Moon & J.E.Han (2023)[27] – Japan
  • Ramalina disparata Krog & Swinscow (1976)[35] – Africa
  • Ramalina dissecta Kashiw. (1988)
  • Ramalina dissimilis Krog (2000)[36] – Tanzania
  • Ramalina dumeticola Krog & Swinscow (1976)[35] – Africa

E

  • Ramalina ecklonii (Spreng.) Meyen & Flot. (1843)
  • Ramalina elegans (Bagl. & Carestia) Jatta (1909)
  • Ramalina erosa Krog (1990)[37]
  • Ramalina erumpens Blanchon, Braggins & Alison Stewart (1996) – Australia, New Zealand
  • Ramalina erythrantha Müll.Arg. (1879)
  • Ramalina escorialensis V.Marcano & A.Morales (1994)[38]
  • Ramalina euphorbiae Vain. (1901)
  • Ramalina europaea Gasparyan, Sipman & Lücking (2017)[39] – Europe
  • Ramalina exiguella Stirt. (1881)
  • Ramalina exilis Asahina (1939)[40] – Japan
  • Ramalina extenuata H.Magn. (1955)

F

  • Ramalina fallax Motyka (1961)
  • Ramalina farinacea (L.) Ach. (1810)
  • Ramalina fasciata Kremp. (1876)
  • Ramalina fastigiata (Pers.) Ach. (1810)
  • Ramalina fastigiatofraxinea Hue (1898)
  • Ramalina faurieana Zahlbr. (1945)
  • Ramalina fecunda Krog & Swinscow (1976)[35] – Africa
  • Ramalina feldmannii Faurel, Ozenda & Schotter (1952)
  • Ramalina fennica Räsänen (1931)
  • Ramalina filicaulis G.N.Stevens (1987)[41] – Australia
  • Ramalina fimbriata Krog & Swinscow (1974)[17] – East Africa
  • Ramalina finkii Zahlbr. (1930)
  • Ramalina fissa (Müll.Arg.) Vain. (1900)[42]
  • Ramalina flabelliformis Asahina (1955)
  • Ramalina flaccidissima Bory (1826)
  • Ramalina flagellifera Vain. (1890)
  • Ramalina fleigiae Gumboski, Eliasaro & R.M.Silveira (2018)[43]
  • Ramalina floccosa Delile (1870)
  • Ramalina foliosa Räsänen (1931)
  • Ramalina fontqueri Maheu (1928)
  • Ramalina fortunatarum Blázquez, Pérez-Vargas & Pérez-Ort. (2024)
  • Ramalina fragilis Aptroot & Bungartz (2007)[34] – Galapagos
  • Ramalina fragilissima Krog (1983)[44]
  • Ramalina fraxinea (L.) Ach. (1810)
  • Ramalina furcellangulida Aptroot (2007)[34] – Galapagos

G

  • Ramalina gallowayi Kashiw., T.H.Nash & K.H.Moon (2007)
  • Ramalina geniculatella Aptroot (2008)[45] – Saint Helena
  • Ramalina glaucescens Kremp. (1881)[46]
  • Ramalina gloriosensis R.Poncet (2021)[47] – Scattered Islands
  • Ramalina gomerana Blázquez, Pérez-Vargas & Pérez-Ort. (2024)
  • Ramalina grappae Sambo (1936)
  • Ramalina grumosa Kashiw. (1987)

H

  • Ramalina hengduanshanensis S.O.Oh & Li S.Wang (2014)[48]
  • Ramalina himalayensis Räsänen (1950)
  • Ramalina hioramii B.de Lesd. (1934)[49]
  • Ramalina hivertiana R.Poncet (2021)[47] – Scattered Islands
  • Ramalina hoehneliana Müll.Arg. (1890)[50]
  • Ramalina hokkaidensis Kashiw. (1986)
  • Ramalina hossei Vain. (1921)[26]
  • Ramalina huei Harm. (1911)[51]
  • Ramalina hypodectodes (Nyl.) Zahlbr. (1930)
  • Ramalina hyrcana Sipman (2011)[52]

I

  • Ramalina implectens Nyl. (1870)[2]
  • Ramalina implexa (Nyl.) Krog (1994)[53]
  • Ramalina inaequalis Nyl. (1870)[2]
  • Ramalina incana Kashiw. (1987)[54]
  • Ramalina inclinata Kashiw., K.H.Moon & M.J.Lai (2006)[55] – Taiwan
  • Ramalina incurvescens H.Magn. (1955)[56]
  • Ramalina indica Fr. (1820)
  • Ramalina inflata (Hook.f. & Taylor) Hook.f. & Taylor (1845)
  • Ramalina inflexa Blanchon, Braggins & Alison Stewart (1996) – New Zealand
  • Ramalina insularum H.Magn. (1955)[56]
  • Ramalina intermedia (Delise ex Nyl.) Nyl. (1873)?[2]
  • Ramalina intermediella Vain. (1921)[57]
  • Ramalina intermixta Gyeln. (1932)
  • Ramalina interponens (Nyl.) Zahlbr. (1930)
  • Ramalina intestiniformis Kashiw. & K.H.Moon (2016)[58] – Korea

J

  • Ramalina jamesii Krog (1990)[37]

K

  • Ramalina kardakovae Oxner (1940)
  • Ramalina kazakhorum Oxner (1940)
  • Ramalina ketner-oostrae Aptroot (2008)[45] – Saint Helena
  • Ramalina krogiae Guissard & Sérus. (2020)[59] – Canary Islands
  • Ramalina kurokawae Kashiw. (1996)[60] – Japan
  • Ramalina kuschei B.de Lesd. (1921)

L

  • Ramalina labiosorediata Gasparyan, Sipman & Lücking (2017)[39] – North America
  • Ramalina lacera (With.) J.R.Laundon (1984)[61]
  • Ramalina lacerata Müll.Arg. (1883)
  • Ramalina laciniata Jatta (1903)
  • Ramalina lafayettii Bory (1828)
  • Ramalina lanceolata Nyl. (1870)[2]
  • Ramalina landroensis Zopf (1906)[62]
  • Ramalina latzelii Zahlbr. (1910)
  • Ramalina lavicola H.Magn. (1945)
  • Ramalina leiodea (Nyl.) Müll.Arg. (1883)
  • Ramalina leptosperma Nyl. (1876)[63]
  • Ramalina leucosticta Taylor (1847)[64]
  • Ramalina litoralis Asahina (1939)
  • Ramalina litorea G.N.Stevens (1986)[16] – Australia
  • Ramalina lojkana Motyka (1961)
  • Ramalina lopezii V.Marcano & A.Morales (1994)[7] – Venezuela
  • Ramalina luciae Molho, Bodo, W.L.Culb. & C.F.Culb. (1981)[65]
  • Ramalina lusitanica H.Magn. (1956)[66]
  • Ramalina luxurians H.Magn. (1942)

M

  • Ramalina maciformis (Delile) Bory (1828)
  • Ramalina macrospora Kashiw. (1987)[54]
  • Ramalina maculata Müll.Arg. (1877)[12]
  • Ramalina maderensis Motyka (1961)
  • Ramalina maegdefraui V.Marcano & A.Morales (2021)[67]
  • Ramalina mahoneyi Quedensley & M.Véliz (2011)
  • Ramalina maritima Krog & Swinscow (1976)[35] – Africa
  • Ramalina marteaui R.Poncet (2021)[47] – Scattered Islands
  • Ramalina mediterranea H.Magn. (1956)
  • Ramalina membranacea Mont. (1839)[68]
  • Ramalina menziesii Taylor (1847)
  • Ramalina meridionalis Blanchon & Bannister (2002)[69] – Australia, New Zealand
  • Ramalina mexicana Vain. (1926)
  • Ramalina meyeri Stein (1888)[70]
  • Ramalina microphylla A.Morales & V.Marcano (1994)[7] – Venezuela
  • Ramalina microspora Kremp. (1876)[71]
  • Ramalina montagnei De Not. (1846)[31]

N

  • Ramalina nervulosa (Müll.Arg.) Abbayes (1952)
  • Ramalina nuda J.Steiner (1899)
  • Ramalina nylanderiana Räsänen (1949)

O

  • Ramalina obtusata (Arnold) Bitter (1901)
  • Ramalina osorioi Kashiw., T.H.Nash & K.H.Moon (2007)[72]

P

  • Ramalina pachyphloea J.Steiner (1904)[19]
  • Ramalina pacifica Asahina (1939)[40] – Japan
  • Ramalina palmiformis Kashiw. & T.H.Nash (1994)[3] – Mexico
  • Ramalina panizzii De Not. (1846)[31]
  • Ramalina papillifera J.Steiner (1907)[73]
  • Ramalina papyracea Blázquez, Pérez-Vargas & Pérez-Ort. (2024)
  • Ramalina paradisensis A.Morales & V.Marcano (1995)[74]
  • Ramalina paraguayensis Gyeln. (1931)[75]
  • Ramalina patelliformis Kashiw. (1987)[54]
  • Ramalina pellucida Taylor (1847)[64]
  • Ramalina perlucens Hue (1899)[76]
  • Ramalina pertenuis H.Magn. (1945)[77]
  • Ramalina pertusa Kashiw. (1985)[78]
  • Ramalina peruviana Ach. (1810)
  • Ramalina pichii Sambo (1941)[79]
  • Ramalina pilulifera Taylor (1847)[64]
  • Ramalina pollinaria (Westr.) Ach. (1810)
  • Ramalina polyforma Aptroot (2007)[34] – Galapagos
  • Ramalina polymorpha (Lilj.) Ach. (1810)
  • Ramalina portosantana Krog (1990)[37]
  • Ramalina portuensis Samp. (1924)[80]
  • Ramalina praetermissa Abbayes (1956)[81]
  • Ramalina prinii B.de Lesd. (1956)
  • Ramalina pseudosekika Asahina (1941)
  • Ramalina psoromica Kashiw. & T.H.Nash (2002)[22] – Mexico
  • Ramalina puberulenta Riefner & Bowler (1994)[82]
  • Ramalina puiggarii Müll.Arg. (1881)[83]
  • Ramalina pusilla Duby (1830)

Q

  • Ramalina qinlingensis S.Y.Guo & L.F.Han (2021)[4] – China
  • Ramalina quercicola Kashiw. & T.H.Nash (2004)

R

  • Ramalina ramificans Seavey & J.Seavey (2017)[84] – Florida
  • Ramalina rectangularis Nyl. (1870)[2]
  • Ramalina reducta Krog & Swinscow (1976)[35] – Africa
  • Ramalina reptans Kashiw., C.W.Sm. & K.H.Moon (2002)[85] – Hawaii
  • Ramalina requienii (De Not.) Jatta (1892)
  • Ramalina retiformis Menzies (1860)
  • Ramalina rigidella Aptroot (2008)[45] – Saint Helena
  • Ramalina riparia Blanchon, Braggins & Alison Stewart (1996) – New Zealand
  • Ramalina rjabuschinskii Savicz (1914)
  • Ramalina roesleri (Schaer.) Nyl. (1870)[2]
  • Ramalina rosacea (A.Massal.) Hepp (1857)
  • Ramalina ryukyuensis Kashiw. & K.H.Moon (2017)[86] – Japan

S

  • Ramalina sabinosana Blázquez, Pérez-Vargas & Pérez-Ort. (2024)[87]
  • Ramalina sampaioana Blázquez, Pérez-Vargas & Pérez-Ort. (2024)[87]
  • Ramalina sanctae-helenae Aptroot (2008)[45] – Saint Helena
  • Ramalina sandwicensis Zahlbr. (1903)[88]
  • Ramalina santanensis V.Marcano & A.Morales (1994)[7] – Venezuela
  • Ramalina sarahae K.Knudsen, Lendemer & Kocourk. (2018)[89] – Channel Islands (California)
  • Ramalina saxicola Zahlbr. ex H.Magn. (1945)
  • Ramalina scoriseda Zahlbr. (1914)[90]
  • Ramalina scrobiculata Müll.Arg. (1885)[91]
  • Ramalina seawardii Aptroot & Sipman (2001)[92] – Hong Kong
  • Ramalina sekika Asahina (1941)
  • Ramalina semicuspidata (Räsänen) Sheard (1978)[93]
  • Ramalina shinanoana Kashiw. (1986)[94]
  • Ramalina sideriza Zahlbr. (1911)[95]
  • Ramalina siliquosa (Huds.) A.L.Sm. (1918)[96]
  • Ramalina similis H.Magn. (1945)
  • Ramalina sinensis Jatta (1902)
  • Ramalina sintenisii Müll.Arg. (1888)[97]
  • Ramalina sonorensis Kashiw. & T.H.Nash (1994)[3] – Mexico
  • Ramalina soraligera H.Magn. (1955)
  • Ramalina sorediantha Nyl. (1870)[2]
  • Ramalina sphaerophora Kashiw. & K.H.Moon (2016)[58] – Korea
  • Ramalina stenospora Müll.Arg. (1877)[12]
  • Ramalina stevensiae Elix (1991)[20] – Norfolk Island
  • Ramalina stoffersii Sipman (2011)[52]
  • Ramalina strepsilis (Ach.) Zahlbr. (1894)[98]
  • Ramalina striatula Nees & Flot. (1835)[99]
  • Ramalina subasperata Nyl. (1876)[63]
  • Ramalina subbreviuscula Asahina (1939)
  • Ramalina subcalcarata V.Marcano & A.Morales (2021)[67]
  • Ramalina subdecipiens J.Steiner (1904)[19]
  • Ramalina subdecumbens Kashiw., K.H.Moon & J.E.Han (2021)[23] – South Korea
  • Ramalina subfarinacea (Nyl. ex Cromb.) Nyl. (1873)[100]
  • Ramalina subfraxinea Nyl. (1870)[2]
  • Ramalina subgeniculata Nyl. (1870)[2]
  • Ramalina subleptocarpha Rundel & Bowler (1976)[101]
  • Ramalina sublitoralis Asahina (1939)
  • Ramalina subpollinaria Nyl. (1870)[2]
  • Ramalina subrotunda Kashiw., C.W.Sm. & K.H.Moon (2002)[85] – Hawaii
  • Ramalina subulata Mont. (1852)[102]
  • Ramalina subvulcania Werner (1948)
  • Ramalina sulcatula Nyl. (1870)[2]

T

  • Ramalina taitensis Nyl. (1870)[2]
  • Ramalina tapperi Krog & Swinscow (1974)[17] – East Africa
  • Ramalina tenaensis V.Marcano & A.Morales (2021)
  • Ramalina tenella Müll.Arg. (1879)[103]
  • Ramalina tenuis Fr. & Tuck. (1858)[104]
  • Ramalina tenuissima V.Marcano & A.Morales (1994)[7] – Venezuela
  • Ramalina thrausta (Ach.) Nyl. (1870)
  • Ramalina thraustoides Vain. (1924)
  • Ramalina throwerae Aptroot & Sipman (2001)[92] – Hong Kong
  • Ramalina timdaliana Krog (1990)[37]
  • Ramalina tingitana Salzm. (1870)[2]
  • Ramalina tovarensis V.Marcano & A.Morales (1994)[7] – Venezuela
  • Ramalina translucida Krog & Swinscow (1974)[17] – East Africa
  • Ramalina tropica G.N.Stevens (1987)[41] – Australia
  • Ramalina tucumanensis Räsänen (1949)
  • Ramalina tumescens Kashiw., K.H.Moon & J.E.Han (2023)[27] – Japan

U

  • Ramalina umbilicata H.Magn. (1945)
  • Ramalina unifolia J.W.Thomson (1990)[105]
  • Ramalina unilateralis F.Wilson (1889)
  • Ramalina usnea (L.) R.Howe (1914)[106]

V

  • Ramalina vareschii V.Marcano & A.Morales (1994)[38]
  • Ramalina venezuelensis V.Marcano & A.Morales (2021)
  • Ramalina verrucosa Hook.f. & Taylor (1844)
  • Ramalina victoriana V.Marcano & L.Castillo (2021)
  • Ramalina vittata Nyl. (1870)[2]
  • Ramalina vizzavonensis Zschacke (1927)
  • Ramalina vogulina Vain. (1928)
  • Ramalina voukii Kušan & Zahlbr. (1933)[107]

W

X

  • Ramalina xalapensis V.Marcano & A.Morales (1994)[38]

Y

  • Ramalina yasudae Räsänen (1940)[109]
  • Ramalina yokotae Kashiw. & K.H.Moon (2017)[86] – Japan
  • Ramalina zollingeri Szatala (1937)[110]

References

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  2. ^ a b c d e f g h i j k l m n o p q r s t Nylander, W. (1870). "Recognitio monographica Ramalinarum" [Monographic revision of the Ramalinas]. Bulletin de la Société Linnéenne de Normandie (in Latin). 4: 101–180.
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  7. ^ a b c d e f g h Marcano, Vicente; Méndez, Antonio Morales; Mendez, Antonio Morales (1994). "New species of Ramalina from Venezuela". The Bryologist. 97 (1): 26–33. doi:10.2307/3243345. JSTOR 3243345.
  8. ^ Sérusiaux, Emmanuël; van den Boom, Pieter; Magain, Nicolas (2021). "Ramalina arsenii, an additional new species in the R. pollinaria group in Western Europe". The Lichenologist. 53 (6): 433–439. doi:10.1017/s0024282921000372.
  9. ^ a b Aptroot, A.; Schumm, F. (2008). "Key to Ramalina species known from Atlantic islands, with two new species from the Azores". Sauteria. 15: 21–57.
  10. ^ Dodge, C.W. (1948). Lichens and lichen parasites. British Australian and New Zealand Antarctic Research Expedition Scientific Reports. B. Vol. 7. pp. 1–276 [217].
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  16. ^ a b c Stevens, G. Nell (1986). "Three new saxicolous Ramalinae from Australia". The Lichenologist. 18 (2): 183–189. doi:10.1017/s0024282986000233.
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