Avion locality

Avion locality
Stratigraphic range: Serpukhovian - Moscovian
Location
RegionPas-de-Calais
CountryFrance
Type section
Named forAvion, Pas-de-Calais

The Avion locality is a carboniferous lagerstätte in France. This lagerstätte is located in a coal slag-heap outside the town of Avion in the region of Pas-de-Calais. Unfortunately, as this slag heap is closed and no new fossils are able to be obtained outside of the paleobiota previously identified. In addition, the coal where the scraps are obtained has not been assigned to any formation.[1]

Paleobiota identified

Arthropods

Arthropods
Genus Species Higher taxon Notes Images
Archaeogerarus A. schubneli Archaeogeraridae (Archaeorthoptera) Placed in a monotypic family[2]
Aviobiella A. garroustei Permobiellidae (Caloneurodea)[3]
Aviobreyeria A. gracilis Breyeriidae (Palaeodictyoptera) Discovered alongside indeterminate immature breyeriids[4]
Aviocladus A. pectinatus Cnemidolestodea (Archaeorthoptera) Indeterminate family, as finer cnemidolestodean relationships are unclear[5]
Aviogramma A. gracilis Caloneurodea Indeterminate family[5]
Aviohapaloptera A. bethouxi Archaeorthoptera Different from all other archaeorthopteran families[5]
Aviologus A. duquesnei Archaeorthoptera Similar to Heterologus[6]
Avionblattinopsis A. oudardi Blattinopsidae (Paoliida) First blattinopsid known from Avion[7]
Avionbrodia A. incompleta Diaphanopterodea?[8]
Avionpaolia A. amansfossila Paoliida Species name translates to “those who love fossils”, recognising the efforts of people who collected fossils from Avion[1]
Avionptera A. communeaui Fatjanopteridae (Plecoptera) Related to the formerly enigmatic genus Fatjanoptera[9]
Avionugonioneura A. jouaulti Nugonioneuridae? (Archaeorthoptera) Most similar to nugonioneurids, but still differs somewhat from them[10]
Avionxixia A. gui Cnemidolestidae (Archaeorthoptera) Similar to the Chinese genus Xixia[2]
Aviorrhyncha A. magnifica Aviorrhynchidae (Hemiptera) Earliest known hemipteran[11]
Avioxyela A. gallica Stem-Hymenoptera (Hymenopterida) Earliest known hymenopteran relative[11]
Beloatta B. duquesnei Eoblattidae (Polyneoptera)[12]
Cacurgus C. avionensis Cacurgidae (Archaeorthoptera) Discovery led to Cacurgidae being moved closer to Orthoptera[13]
Carbonatura C. oudardi Carbonaturidae (stem group Auchenorrhyncha)[14]
Carbopsyllidium C. minutum Permopsyllidiidae (Hemiptera)[14]
Enigmaptera E. magnifica Paneodonatoptera (Odonatoptera) One of numerous convergently damselfly-like odonatopterans[15]
Fusiogramma F. minuta Caloneurodea (Archaeorthoptera) A relatively small archaeorthopteran[1]
Gallotupus G. oudardi Meganeuridae One of the first insects described from Avion[16]
Gulou G. oudardi Plecoptera Shares a genus with a fossil from the Yanghugou Formation[9]
Homoiopteridae indet. Unapplicable Paleodictyoptera Known from a base of a ~20 cm-long wing[5]
Jacquesoudardia J. magnifica Protozygoptera Bears convergently damselfly-like wings[5]
Paolia P. sp Paoliida Known from a fragmentary wing[5]
Piesbergopterum P. avionensis Cnemidolestodea (Archaeorthoptera) Also known from Piesburg quarry as its name suggests[17]
Spilaptera S. splendens Spilapteridae Named after the fossil’s exquisite preservation[5]
Theiatitan T. azari Titanoptera Earliest known titanopteran, may have communicated via wing-produced light flashes or crepitation (crackling sounds produced from wingbeats in some orthopterans)[18]
Westphalomerope W. maryvonneae Protomeropidae (Panorpida) Earliest holometabolan insect discovered (when it was described)[19]
Westphalopsocus W. pumilio Westphalopsocidae (Psocodea) Earliest known psocodean[11]
Westphalothripides W. oudardi Westphalothripidesidae (Stem-Thysanoptera) Earliest known thrips[20]

References

  1. ^ a b c Boderau, Mathieu; Roques, Patrick; Nel, André (2024). "New paleopteran and polyneopteran insects from the Carboniferous of Northern France". Acta Palaeontologica Polonica. 69. doi:10.4202/app.01189.2024.
  2. ^ a b Nel, André; Roques, Patrick (2021). "A new family and two new genera from Avion, Northern France, confirm the Moscovian, late Carboniferous diversity of the insect superorder Archaeorthoptera". Acta Palaeontologica Polonica. 66 (4): 879–884. doi:10.4202/app.00940.2021.
  3. ^ Nel, André; Roques, Patrick (31 August 2021). "The second Caloneurodea from the Moscovian of Avion, France (Insecta, Archaeorthoptera)". Palaeoentomology. 4 (4). doi:10.11646/palaeoentomology.4.4.5.
  4. ^ Prokop, Jakub; Tippeltová, Zuzana; Roques, Patrick; Nel, André (2013). "A new genus and species of Breyeriidae and wings of immature stages from the Upper Carboniferous, Nord-Pas-de-Calais, France (Insecta: Palaeodictyoptera)". Insect Systematics & Evolution. 44 (2): 117–128. doi:10.1163/1876312X-44032098.
  5. ^ a b c d e f g Prokop, Jakub; Roques, Patrick; Nel, André (3 April 2014). "New non-holometabolous insects from the Pennsylvanian of the Avion locality in Pas-de-Calais, France (Insecta: 'Exopterygota')". Alcheringa: An Australasian Journal of Palaeontology. 38 (2): 155–169. doi:10.1080/03115518.2014.848620.
  6. ^ Coty, David; HáVa, Jiří; Prokop, Jakub; Roques, Patrick (29 October 2014). "New archaeorthopteran insects from the Late Carboniferous of the Nord and Pas-de-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera)". Zootaxa. 3878 (5). doi:10.11646/zootaxa.3878.5.4.
  7. ^ Quispe, Laura; Roques, Patrick; Garrouste, Romain; Nel, André (4 March 2022). "Carboniferous Blattinopsidae: revision of Klebsiella and new genus and species from Avion (Insecta, Paoliida)". Historical Biology. 34 (3): 383–389. doi:10.1080/08912963.2021.1916817.
  8. ^ Schubnel, Thomas; Roques, Patrick; Oudard, Jacques; Garrouste, Romain; Nel, André (26 February 2020). "A new paleopteran genus and species from the late Carboniferous of Avion France, (Insecta: Palaeodictyopterida: ?Diaphanopterodea)". Palaeoentomology. 3 (1): 032–035. doi:10.11646/PALAEOENTOMOLOGY.3.1.3.
  9. ^ a b Schubnel, Thomas; Perdu, Lubin; Roques, Patrick; Garrouste, Romain; Nel, André (3 July 2019). "Two new stem-stoneflies discovered in the Pennsylvanian Avion locality, Pas-de-Calais, France (Insecta: 'Exopterygota')". Alcheringa: An Australasian Journal of Palaeontology. 43 (3): 430–435. doi:10.1080/03115518.2019.1569159.
  10. ^ Nel, André; Roques, Patrick (2 September 2022). "A new strange Archaeorthoptera from the Moscovian of Avion (France) (Insecta, Polyneoptera)". Historical Biology. 34 (9): 1769–1771. doi:10.1080/08912963.2021.1978082.
  11. ^ a b c Nel, André; Roques, Patrick; Nel, Patricia; Prokin, Alexander A.; Bourgoin, Thierry; Prokop, Jakub; Szwedo, Jacek; Azar, Dany; Desutter-Grandcolas, Laure; Wappler, Torsten; Garrouste, Romain; Coty, David; Huang, Diying; Engel, Michael S.; Kirejtshuk, Alexander G. (14 November 2013). "The earliest known holometabolous insects". Nature. 503 (7475): 257–261. doi:10.1038/nature12629.
  12. ^ Nel, André; Garrouste, Romain; Roques, Patrick (21 December 2020). "The first representative of the archaeorthopteran family Eoblattidae in the Konservat Lagerstätte of Avion, France (Insecta: Polyneoptera)". Palaeoentomology. 3 (6): 552–555. doi:10.11646/palaeoentomology.3.6.3.
  13. ^ Schubnel, Thomas; Roberts, Dawn; Roques, Patrick; Garrouste, Romain; Desutter-Grandcolas, Laure; Nel, André (18 March 2020). "Moscovian fossils shed light on the enigmatic polyneopteran families Cacurgidae and Eoblattidae (Insecta: 'Eoblattida', Archaeorthoptera)". Journal of Systematic Palaeontology. 18 (6): 499–511. doi:10.1080/14772019.2019.1627595.
  14. ^ a b Boderau, M.; Schubnel, T.; Fu, Y.; Mottequin, B.; Robin, N.; Roques, P.; Kundura, J.-P.; Garrouste, R.; Nel, A. (2026). "Carboniferous fossils enlighten the systematics and evolution of Hemiptera". Papers in Palaeontology. 12 (2) e70073. doi:10.1002/spp2.70073.
  15. ^ Nel, André; Roques, Patrick; Prokop, Jakub; Garrouste, Romain (3 April 2019). "A new, extraordinary 'damselfly-like' Odonatoptera from the Pennsylvanian of the Avion locality in Pas-de-Calais, France (Insecta: 'Exopterygota')". Alcheringa: An Australasian Journal of Palaeontology. 43 (2): 241–245. doi:10.1080/03115518.2018.1489561.
  16. ^ Roques, Patrick; Garrouste, Romain; Nel, Andre (2008). "A new griffenfly genus from the Late Carboniferous of northern France (Odonatoptera: Meganeuridae)". Insect Systematics & Evolution. 39 (2): 231–239. doi:10.1163/187631208794760894.
  17. ^ Nel, André; Roques, Patrick (31 August 2021). "The second species of the cnemidolestid genus Piesbergopterum from the Upper Carboniferous of Avion, Northern France (Archaeorthoptera: Cnemidolestidae)". Palaeoentomology. 4 (4): 323–325. doi:10.11646/PALAEOENTOMOLOGY.4.4.6.
  18. ^ Schubnel, Thomas; Legendre, Frédéric; Roques, Patrick; Garrouste, Romain; Cornette, Raphaël; Perreau, Michel; Perreau, Naïl; Desutter-Grandcolas, Laure; Nel, André (8 July 2021). "Sound vs. light: wing-based communication in Carboniferous insects". Communications Biology. 4 (1). doi:10.1038/s42003-021-02281-0. PMC 8266802.
  19. ^ Nel, André; Roques, Patrick; Nel, Patricia; Prokop, Jakub; Steyer, J. Sébastien (January 2007). "The earliest holometabolous insect from the Carboniferous: a "crucial" innovation with delayed success (Insecta Protomeropina Protomeropidae)". Annales de la Société entomologique de France. New Series. 43 (3): 349–355. doi:10.1080/00379271.2007.10697531.
  20. ^ Nel, Patricia; Azar, Dany; Prokop, Jakub; Roques, Patrick; Hodebert, Gilbert; Nel, André (June 2012). "From Carboniferous to Recent: wing venation enlightens evolution of thysanopteran lineage". Journal of Systematic Palaeontology. 10 (2): 385–399. doi:10.1080/14772019.2011.598578.
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